16 de setembro de 2021

Range and population observations on some of the more commonly reported cynipid galls in the Eastern US.

Was bored at work today waiting for a bioinformatic analysis to run, so decided to make some notes on the distribution of the more commonly reported oak gall wasps. Also included here are some remarks on the local populations of a few species that I have observed over several years.

Acraspis erinacei – generally more northern, present but less common in the south. In Ohio, one of the most common and most consistent year-to-year.
Acraspis quercushirta – Centered on Appalachians and East. Based on the map, looks to be most common on Q. montana, compared to its other hosts.
Acraspis macrocarpae – Northern distribution, also on Q. gambeli in the Rockies, has extended beyond the native range of its host to cultivated trees in Alberta, present in both Calgary and Edmonton as well as Missoula, Montana. Absent from the south.
Acraspis pezomachiodes – more northern as well. More compact distribution compared with A. erinacei, with which it is found often in Ohio. Very common in Ohio and consistent year-over-year populations.
Acraspis prinoides – interesting Midwest centered distribution, more reported on Q. muehlenbergii than Q. montana? Given the host specificity of Acraspis species and Andrew Hipp’s recent oak phylogeny, we should rear some of these to confirm they are the same species. Uncommon in Ohio, very rare to find a tree with more than a couple.
Amphibolips acuminata - Mid-Atlantic centered distribution, with scattered observations in the southern states. Commonly found on Q. imbricaria in swamps in NE Ohio.
Amphibolips citriformis – southern distribution consistent with the ranges of its hosts.
Amphibolips confluenta – widespread, eastern NA. Has exceeded natural range of host – credible observations in Lubbock TX and Albequerque NM on cultivated trees.
Amphibolips cookii – more common in the north, esp the NE. Very common in Ohio, rare to find a group of Q. rubra without one. Consistent population year-over-year.
Amphibolips gainesi - Texas-centered distribution. Its hosts are throughout the SE, so may be some sampling bias given the great gall hunters that we have in TX.
Amphibolips nublipennis – more common north and east. Not a common gall in NE Ohio, I have seen it most frequently on Q. imbricaria compared to its other hosts.
Amphibolips quercuscinerea – almost exclusively Florida. Supposedly hosted by Q. incana, which has a wider distribution, but is probably undersurveyed.
Amphibolips quercusinanis – widely distributed but not as far south as A. confluenta. Very common in Ohio and consistant year-over-year. In my area much more common than A. confluenta.
Amphibolips quercusjuglans – widely distributed in eastern NA, has exceeded natural range of hosts in at least one example – Abilene TX. Anecdotally, seems to have population fluctuations locally here in Ohio, a mega crop of these in some years and few and far between in others, related to mast years in their oak hosts?
Amphibolips quercusostensackenii – more common in north. Very common and consistent year-over-year in NE Ohio. On nearly every pin oak.
Amphibolips quercusracemaria – Southern coastal plain/piedmont. Especially along gulf coast. Hotspot in DFW.
Amphibolips quercusrugosa – Midwest/midatlantic-centered distribution, although might be some sampling bias bc this one is not particularly charismatic. In Ohio most common on Q. imbricaria on wetland sites.
Andricus aciculatus – Southern distribution. Were extremely common at one of the Q. lyrata sites that I visited in S. Illinois. Never observed on Q. stellata in Ohio.
Andricus biconicus – Scattered throughout eastern US on Q. stellata. Seems to be more common in Midwestern Q. stellata populations than southern ones, based on the relatively few reports from the DFW area. Very common on Ohio post oaks.
Andricus chinquapin – mid-Atlantic centered distribution, but very possible that there is sampling bias, given how subtle these are.
Andricus coronus – seems to be more common east of the Appalachians, scattered elsewhere. I have found at only one random site in NE Ohio, and have looked at 1000s of pin oaks.
Andricus dimorphus – Distribution centered on Midwest, somewhat mirrors the distribution of its most common host, Q. macrocarpa. Promiscuous, host-wise, and has consistent year-over-year populations.
Andricus ignotus – rare in SE, scattered elsewhere. Fairly common on bur oak in Ohio.
Andricus nigricens – Midwest-NE centered distribution. Extremely common on Q. bicolor in NE Ohio to the point of which I use it to ID the tree.
Andricus pattoni – more southerly distribution, but found wherever Q. stellata are. Very common on Q. stellata in Ohio.
Andricus quercusflocci – even distribution in E. NA. Seems to skew slightly more southern than other Q. alba galls given that is has only been observed once in Michigan, Minnesota or Ontario. Common in NE Ohio, though not as much as either of the Acraspis galls on Q. alba.
Andricus quercusfoliatus – Southern coastal plain and Texas.
Andricus quercusformosus – Eastern coastal plain-centered distribution. On my spring trip, this year was locally very common on Q.incana and Q. laevis in some Longleaf pine forests… seems to be rare elsewhere.
Andricus quercusfrondosus – Northern distribution (DFW hotspot). In Ohio, definitely more common on Q. macrocarpa than its other hosts and thus more common in the W. half of the state where that species is more concentrated.
Andricus quercuslanigera – Mostly southern coastal plain species, but has followed Q. virginiana wherever that species has been planted even to California.
Andricus quercuspeticola – widespread E. NA. Host promiscuous.
Andricus quercusstrobilanus – Midwest/midatlantic centered distribution, hot spot in Chicago (large bur oak concentration?). I have seen these by far most frequently on Q. macrocarpa compared to its other hosts.
Andricus quercusutriculus – Currently midwest/mid-atlantic centered distribution. But definitely suffers from sampling bias. Very subtle species.
Andricus weldi – Midwest/midatlantic. I suspect this is more common than reported bc it is somewhat subtle. Very common in NE. Ohio, esp on mature trees.
Atrusca quercuscentricola –An odd Missouri/Arkansas centered distribution – conspicuously rare from the very well reported DFW area. Has some color variation across the range, with the reds that are apparent in the Missouri/Arkansas populations not encountered as much farther east. Very common on Q. stellata in Ohio, esp at the darby plains sites.
Belonocnema kinseyi - Texas-centered distribution.
Belonocnema treatae – Florida-centered distribution.
Callirhytis clavula – more common north and east. Very common on Q. alba in NE Ohio, have not encountered elsewhere in Ohio.
Callirhytis favosa – Northern distribution, have noticed population dynamics in this species, common some years and rare in others.
Callirhytis furva – eastern US, in NE Ohio undergos extreme population fluctuations, anecdotally have noticed an every other year pattern, with this sometimes being the single most common gall and being much lower in alternate years – one of the most common galls in NE Ohio in summer 2020 and have only seen a couple times 2021.
Callirhytis gallaestriatae -- Seems to be northerly distributed but could be sampling bias given that these are extremely subtle. Fairly common in Ohio, very early in spring (before leaf out).
Callirhytis infuscata – interesting distribution mostly on a diagonal line from Toronto ON, to DFW, probably sampling bias. Not common in Ohio.
Callirhytis lanata – more common north and east. Very common in NE Ohio, hard to find a Q. rubra without one.
Callirhytis piperoides – northeast centered distribution. Not common in Ohio.
Callirhytis quercusbatatoides – S. Coastal plain and Texas.
Callirhytis quercuscornigera – Midwest/north Atlantic centered distribution. Probably most common on Q. imbricaria in NE Ohio, but fairly host-promiscuous.
Callirhytis quercusfutilis – Mid-atlantic centered distribution. Common in NE Ohio and most frequently seen on Q. alba and Q. bicolor.
Callirhytis quercusgemmaria – scattered E NA. Not common in NE Ohio, but subtle and so probably underreported.
Callirhytis quercusoperator – common on the east coast, scattered elsewhere (rare in NE Ohio).
Callirhytis quercusventricosa – Scattered distribution E NA. Very local populations in my experience. Anecdotally, seems to like less disturbed spots than many other species. Massive host list but I have only observed on Q. imbricaria in NE Ohio. Unlike other Q. imbricaria galls, I have found this one more frequenly on dry sites.
Callirhytis seminator – Widely distributed across E NA but more common East of the Appalachians (uncommon in NE Ohio). So showy as to be relatively over-reported.
Callirhytis vaccinii – Exclusively in Texas.
Disholcaspis cinerosa – Texas-centered distribution.
Disholcaspis globosa – More common Appalachians and east. Rare in Ohio.
Disholcaspis pruniformis – Texas-centered distribution.
Disholcaspis quercusglobulus – Wide distribution in Eastern NA. Very common and consistent populations year-over-year in NE Ohio.
Disholcaspis quercusmama – Somewhat northern distribution, where its two most frequent hosts (Q. macrocarpa and Q. bicolor) are more common.
Disholcaspis quercusvirens – E coastal plain and Texas, especially common in Florida.
Disholcaspis spongiosa – Southern distribution
Dryocosmus floridensis – Scattered distribution in E. US. Very common in NE Ohio on Q. imbricaria.
Dryocosmus quercuspalustris – more common north and east, but a hot spot in DFW as well. Extremely common in NE Ohio, and consistant year-over-year.
Kokkocynips decidua – NE centered distribution, Minnesota also a hot spot. Rare in Ohio (never personally observed).
Kokkocynips difficilis – Piedmont/coastal plain species. One of the few that looks to be more in the piedmont than the coastal plain, but could just be sampling bias.
Kokkocynips imbricariae – wide distribution E. NA, in NE Ohio most common on Q. rubra, but have seen on other host species as well.
Kokkocynips rileyi – more common in parts North and East, but a hot spot in DFW... might be some samping bias given that these are fairly tiny. In NE Ohio most common on Q. imbricaria and velutina, but have seen on most of the red oak group. Might have a couple of flushes in a year, seem to see fresh ones in June and again in late August/September.
Melikaiella tumifica – NE centered distribution. Present in Ohio, but not super common, easy to overlook
Neuroterus quercusirregularis - Seems like a trend to be more southern in distribution (but needs more sampling).
Neuroterus quercusbatatus – NE/Mid-Atlantic centered distribution. Scattered in NE. Ohio.
Neuroterus quercusvarrucarum – Scattered distribution in E. NA with hot spots (DFW, DC). Have noticed population fluxutions in this species, some years extremely common, some years rare in NE Ohio. On Ohio States main campus in Columbus was one of the single most common galls.
Neuroterus saltarius – Northern distribution, certainly more common than reported.
Neuroterus tantalus – Northern distribution, more common than reported. On nearly every Q. alba in NE Ohio.
Neuroterus umbilicatus – Midwest/mid-atlantic centered distribution, could be sampling bias as these are subtle. Easy to find, but not overly common in NE Ohio. Some year-to-year flux in populations.
Neuroterus vesicula – Midwest/mid-atlantic centered at the moment, but almost certainly subject to sampling bias.
Philonix nigra – more northern distribution. Some flux in populations, 2021 a low year in my experience.
Phylloteras nigrum – Midwest/midatlantic centered distribution, certainly some sampling bias here as they occur at a season (late September-October) when fewer people are looking. Not hard to find in NE Ohio, but not common either.
Phylloteras poculum – Midwest/mid-atlantic centered distribution. Common in NE Ohio but not as much as the Acraspis galls on Q. alba. Definitely more common on older trees in my experience.

Phylloteras vollutellae – Northern distribution, definitely more common in areas where host is common (Present in W. Ohio, have not found in NE Ohio). Seems to be very common in Chicagoland where Q. macrocarpa is king.
Trigonaspis polita – Disjunct peaks in population in Texas and Florida, scattered observations up the E coast.
Trigonaspis quercusforticorne – Far northern distribution. Chicagoland and Massachusetts the farthest south observations. Majority of observations in Canada.
Zopheroteras compressum – more northern distribution, although sampling bias definitely probable. Another late season gall.
Zapatella quercusphellos – NE/Mid-Atlantic centered distribution. Infrequent in NE Ohio, most common on Q imbricaria here.
Zopheroteras guttatum – more common North and East, with a hot spot in DFW, more common than reported (tiny size). Flucuates in population, in sync with C. furva.
Zopheroteras sphaerula – NE/midwest centered distribution, sampling bias a possibility as this is another tiny one.

Overall trends and questions
Many Q. alba galls (with the notable exception of DQG, A. quercusflocci and C. seminator) are more common in the north, Q alba rarer or more scattered in the South?
Several red oak galls are focused more north and east as well, again with a few notable exceptions. Does this just represent sampling error, is it a factor of tree density or something else?
Post oak galls tend to be southerly distributed, which makes perfect sense, but many are underreported in areas other than Texas… if you live in an area with post oaks go out and take a look. I think one of the problems here is that in areas other than TX/OK post oaks are not common in cultivation, and thus you need to go to rural areas to find them.
What drives population fluctuations in some species? Is it just simply that they emerge every other year? Do these fluctuations represent an adaptation to reduce parasitoid pressure?
How are some species able to expand their range beyond the range of their host while others stay quite local? Are some species more adaptable to wind distribution or hitching a ride on nursery plants?

Needs
Surveying of common but less charismatic species, particularly small bud galls and integral leaf galls.
Surveying of the less populated southern states - Northern Louisiana across through South Carolina.
Surveying of non-summer species - spring and fall species are less well surveyed.
Surveying of less common oak species that may be strong hosts for some of these "rare" species

Anyone with any thoughts on any of these species? Do your local populations match these observations or contradict them?

Publicado em 16 de setembro de 2021, 08:22 PM por calconey calconey | 4 comentários | Deixar um comentário

29 de junho de 2021

Ohio Gall Checklist

Ohio Gall Checklist
A draft of a gall checklist for Ohio, sorry about inconsistent common/scientific name usage, capitalization and loose host associations. Will edit to correct these details in the future. List contains mostly described species, will add undescribed species in the future.

Comments represent my personal experience hunting galls in Ohio over the last ~5 years (seriously in the past 3)

Gall Wasps
Acraspis erinacei – white oak – widespread, common – very late spring-fall, leaves
Acraspis gemula – white oak – uncommon – spring, buds
Acraspis macrocarpae – bur oak – widespread, common – very late spring-fall, leaves
Acraspis pezomachoides – white oak – widespread, common – very late spring-fall, leaves
Acraspis prinoides – chestnut oak – uncommon – summer-fall, leaves
Acraspis quercushirta – Chestnut/chinkapin oak - ? never personally observed
Acraspis villosa – bur oak – widespread, common – summer-fall, leaves
Amphibolips acuminata – shingle oak – uncommon – late spring, buds/stems
Amphibolips femoratus – red oaks - ? (not personally observed)
Amphibolips globus – pin oak – rare – late spring-early summer, buds
Amphibolips confluenta – Red oaks – widespread, common – late spring, buds
Amphibolips cookii – red oaks – widespread, uncommon – late summer-fall, buds
Amphibolips ellipsodalis – black oak, shingle oak – uncommon – spring, buds
Andricus nigricens – swamp white oak – uncommon – mid-summer, leaves
Amphibolips nublipennis – red oaks – uncommon – late spring, leaves
Amphibolips quercusinanis – red oaks – widespread, common – spring, leaves
Amphibolips quercusjuglans – red oaks – widespread, uncommon – summer, acorns, on ground under trees
Amphibolips quercusoostensackii – red oaks – widespread, common – spring, dried galls through summer/fall, leaves
Amphibolips quercusrugosa – shingle oak – locally common esp. in NE Ohio – spring, leaves
Andricus apiarium -q alba - rare - fall, leaves
Andricus biconicus – post oak – locally common on host, Madison county presumably south to Adams county – late summer/fall - buds
Andricus capillatus – white oak – uncommon? – late summer, leaves
Andricus chinquapin – white oak – uncommon – spring, leaves
Andricus comata – white oak – common? – spring, leaves
Andricus coronus – pin oak – very rare – early spring, buds
Andricus dimorphis – white oaks – widespread, common – summer/fall, leaves
Andricus foliaformis – bur oak – rare – spring, leaves
Andricus frondosus – bur oak – uncommon, more common as you move west and south, spring/summer, buds
Andricus incertus – bur oak, swamp white oak – rare – summer, acorns
Andricus ignotus – bur oak – uncommon – summer, leaves
Andricus quercusflocci – white oak – widespread, uncommon – summer/fall, leaves
Andricus quercuspeticola – white oak – widespread, somewhat common – spring/summer, leaves
Andricus quercusstrobiloides – bur oak, white oak – widespread, uncommon – summer/fall, buds/stems
Andricus mamilaformis – white oak – uncommon-rare – late summer/fall, buds
Andricus pattoni – post oak – Adams county, rare – late summer/fall, leaves
Andricus pisiformis – white oak – uncommon – early spring, buds
Andricus quercusutriculus – white oak – uncommon – spring, leaves
Andricus robustus – bur oak, post oak – rare, Crawford county to parts south and west, more common on post oak – late summer/fall, leaves
Andricus stropus – post oak – locally uncommon, Madison county south to Adams county – summer/fall, buds
Andricus weldi - white oak – locally common – summer/fall, petioles/leaf-base
Atrusca carolina – post oak – rare, Adams county – summer/fall, leaves
Atrusca quercuscentricola – post oak – locally common Madison county south to Adams county – late summer/fall, leaves
Atrusca unica – post oak – uncommon, Madison county south to Adams county – late summer/fall, leaves
Callirhytis balanacea – pin oak - ? late spring/summer
Callirhytis bipapillata – shingle oak – rare? (not personally seen)
Callirhytis clavula – white oak – uncommon – fresh in late spring/summer, galls persist throughout year, stems
Callirhytis confusa – shingle oak – rare – spring, leaves
Callirhytis excavata – red oaks – rare - stems
Callirhytis exigua – white oak – rare, only observed in Washington county – spring, flowers
Callirhytis flavipes – bur oak – widespread, uncommon – spring, leaves
Callirhytis furva – pin oak and other red oaks – widespread, common – summer/fall, leaves
Callirhytis gallaestriatae – pin oak – uncommon – early spring, stems
Callirhytis gemmiformis – white oak – rare – summer/fall, stems
Callirhytis glandulus - white oaks -? - summer, fruits
Callirhytis infuscata – red oaks – uncommon – late summer/fall, leaves
Callirhytis lanata – red oak – widespread, common – summer/fall, leaves
Callirhytis nigrae – shingle oak – rare? – late spring?, leaves
Callirhytis quercuscornigera – red oaks – widespread, common – fresh in spring, persist through the year, stems
Callirhytis quercusfutilis – white oak, swamp white oak – widespread, common – spring, leaves
Callirhytis quercusgemmaria – red oaks – rare – late spring, stems
Callirhytis quercusoperator – red oak – rare – spring, flowers
Callirhytis quercuspunctata – red oaks - ? - stems
Callirhytis quercusventricosa – red oaks , esp shingle oak – rare – spring, stems
Callirhytis parva – shingle oak – rare? – spring, flowers
Callirhytis pedunculata – red oaks - rare? – spring, leaves
Callirhytis piperoides – red oaks – ? never personally observed – late summer/fall, leaves
Callirhytis pulchra – red oak – rare? – spring, flowers
Callirhytis scitula – shingle oak - ? – late spring/summer, stems
Callirhytis seminator – white oak – widespread, uncommon – spring, dried galls persist through summer, stems
Diastrophus cuscutaeformis – rubus – rare (never personally seen) - ?, stems
Diastrophus nebulosus – rubus – uncommon - ? persist through year, stems
Diplolepis bicolor – native rose – rare? Never personally observed – late spring, stems/leaves
Diplolepis dichlocera – native rose – rare? – ? persist through year, stems
Diplolepis nervosa – native rose – rare – spring, leaves
Diplolepis polita – native rose – rare – late spring, leaves/stems
Diploleips rosae – cultivated European roses – becoming rare – spring/summer, stems
Diplolepis rosaefolii – Rosa sp. - uncommon - summer
Disholcaspis globosa - chestnut oak - ? probably more common in SE Ohio – summer/fall, lower stems close to ground
Disholcaspis pruniformis – post oak – Rare? Presumably Madison county south to Adams county – summer, stems
Disholcaspis quercusglobulus – white oak, chestnut oak – widespread, common – summer/fall, persisting, stems
Disholcaspis quercusmama – bur oak, swamp white oak – widespread, common – summer/fall, persisting, stems
Dryocosmus albidus – red oaks – locally uncommon – early summer, leaves
Dryocosmus cinerea – red oaks – uncommon – spring, leaves
Dryocosmus floridensis – red oaks, especially shingle oak – locally common – spring, stems
Dryocosmus kuriphilus – native and cultivated chestnuts – widespread, common – spring/summer, leaves
Dryocosmus quercusnotha - red oaks – uncommon – spring, leaves
Dryocosmus quercuspalustris – pin oak and other red oaks – widespread, common – spring, leaves
Hemadas nubilipennis – blueberry – common in NE Ohio surrounding bogs/fens – summer/fall, stems
Kokkocynips decidua – red oaks – rare? (never personally seen) – late summer/fall, leaves
Kokkocynips imbricariae – red oaks – widespread, uncommon – summer/fall, stems
Kokkocynips rileyi – shingle oak, black oak other red oaks – widespread, common – late spring-fall, leaves
Liposthenes glechomae – Ground Ivy (Glechoma) – widespread, uncommon – spring/summer, leaves
Loxaulus mammula – white oak – uncommon – late spring, stems
Melikaiella sp (formerly Callrhytis quercusmodesta) – red oaks – widespread, uncommon – spring, leaves
Melikaiella tumifica – red oak – uncommon – spring, leaves
Neuroterus clarkeae – white oak – uncommon? – spring, leaves
Neuroterus laurifoliae - shingle oak - ? – late summer/fall, leaves
Neuroterus minutus – white oak – uncommon – spring, buds/leaves
Neuroterus perminimus – white oak – rare? – summer, leaves
Neuroterus quercusbatatus – white oak – uncommon - ?, summer, stems
Neuroterus quercusirregularis – white oaks – uncommon – spring, leaves
Neuroterus quercusverrucarum – bur oak – widespread, common – summer/fall, leaves
Neuroterus saltarius – bur oak, swamp white oak – widespread, common – late spring, persisting into summer, leaves
Neuroterus tantalus – white oak – widespread, common – late spring, persisting into summer, leaves
Neuroterus umbilicatus – white oak – widespread, uncommon - summer/fall, leaves
Neuroterus vesicula – white oak – widespread, common – very early spring, buds
Philonix nigra – white oaks – widespread, common – very late spring/fall, leaves
Phylloteras nigrum – white oaks – uncommon – fall, leaves
Phylloteras poculum – white oak – widespread, uncommon – summer/fall, leaves
Phylloteras prinum - chestnut oak - rare, summer, probably more common in parts east and south, given the distribution of the host
Phylloteras rubinum – white oak – uncommon – fall, leaves
Phylloteras volutellae - bur oak – uncommon, western ohio (probably Crawford county and west) – summer/fall, leaves
Zapatella quercusphellos – shingle oak – uncommon – spring, stems
Zopheroteras compressum – red oak – widespread, uncommon – late summer/fall, leaves
Zopheroteras hubbardi – red oaks esp black oak – uncommon – late summer/fall, leaves
Zopheroteras guttatum – red oaks esp pin oak, widespread, common – summer/fall, leaves
Zopheroteras sphaerula – red oaks, esp q. rubra – widespread, common – summer, leaves

Gall Midges
Acericecis ocellaris – maples, esp red maple – widespread, common – spring, leaf spot
Ampelomyia viticola – grapes – rare – late spring/summer, leaves
Ampelomyia vitiscoryloides – grapes – uncommon – summer, stems
Ampelomyia vitispommum – grapes – rare – summer, stems
Anthodiplosis eutrochii – joe pye weed – scattered, locally common – summer, flower buds
Asphondylia eupatorii – Ageratina – widespread, common – summer/fall, stems
Asphondylia helianthiglobulus – sunflower – common in W. Ohio, not seen in NE – summer/fall, stems
Asphondylia monacha – Solidago – rare? – summer, terminal bud
Asphondylia pseudorosa – Euthamia – rare? – summer, buds
Asphondylia rudbeckiaeconspicua – rudbeckia lacinata – widespread, uncommon – summer, terminal bud
Asphondylia solidaginis – solidago – widespread, common – late spring/fall, leaves
Asteromyia carbonifera – Solidago – widespread, common – late spring/fall, leaves
Asteromyia euthamiae – Euthamia – widespread, common – late spring/fall leaves
Asteromyia modesta – various Asteraceae, commonly on fleabane – uncommon – summer, leaves
Blaesodiplosis crataegifolia – hawthorn (esp crus-galli type) – locally common, more common in NE Ohio – spring, leaves
Blaesodiplosis venae – hawthorn – uncommon – spring, leaves
Caryadiplosis venicola – carya – widespread, common – late spring/fall, leaves
Caryomyia aggregata – carya – uncommon – summer/fall, leaves
Caryomyia ansericollum – bitternut hickory – common – summer/fall, leaves
Caryomyia antennata – carya – uncommon – summer, leaves
Caryomyia asteris – carya – rare – summer, leaves
Caryomyia biretta – carya – locally common – summer, leaves
Caryomyia caryae – carya – widespread, common – summer, leaves
Caryomyia caryaecola – carya – locally common – summer, leaves
Caryomyia conoidea – carya – uncommon – summer, leaves
Caryomyia deflexipili – carya – rare – summer, leaves
Caryomyia eumaris - carya - rare - summer, leaves
Caryomyia flaticrustum – carya – uncommon – summer, leaves
Caryomyia glauciglobus – carya – uncommon-rare – summer, leaves
Caryomyia hirtidolium – carya – rare – summer, leaves
Caryomyia holotricha – carya – rare? – summer, leaves
Caryomyia lenta – carya – uncommon – summer, leaves
Caryomyia inanis – carya – rare – summer, leaves
Caryomyia inclinata – carya – uncommon – summer, leaves
Caryomyia inflata – carya - ? – summer, leaves
Caryomyia levicrustum – carya - ? – summer, leaves
Caryomyia leviglobus carya – rare – summer, leaves
Caryomyia mariginata – carya – uncommon – summer, leaves
Caryomyia persicoides – carya – uncommon – summer, leaves
Caryomyia procumbens – carya – widespread, uncommon – summer, leaves
Caryomyia purpurea – carya – uncommon – summer, leaves
Caryomyia recurvata – carya – uncommon – summer, leaves
Caryomyia sanguinolenta – carya – uncommon – summer, leaves
Caryomyia spiniglobulus – carya – rare – summer, leaves
Caryomyia spinulosa – bitternut – uncommon – summer, leaves
Caryomyia stellata – carya – widespread, uncommon – summer, leaves
Caryomyia striolata – carya – uncommon? – spring, leaves
Caryomyia subulata – carya – uncommon – summer, leaves
Caryomyia supina – carya – uncommon – summer, leaves
Caryomyia thompsoni – widespread, common – summer, leaves
Caryomyia tuberculata – carya – uncommon – summer, leaves
Caryomyia tuberidolium – carya – uncommon – summer, leaves
Caryomyia tubicola – Carya – widespread, common – summer, leaves
Caryomyia tumida – carya - ? – summer, leaves
Caryomyia viscidolium – carya – widespread, common – summer, leaves
Celticecis acuminata – celtis – rare, SW Ohio – late spring/summer, leaves
Celticecis celtiphylla – celtis – uncommon – late spring/summer, leaves
Celticecis conica – celtis – western ohio, uncommon – late spring/summer, leaves
Celticecis corunata – celtis – scattered, uncommon – late spring/summer, leaves
Celticecis expulsa – celtis – uncommon to rare (never personally seen) – late spring, leaves
Celticecis ovata – celtis – widespread, uncommon – late spring/summer, leaves
Celticecis oviformis – celtis – rare? – late spring/summer, leaves
Celticecis pyriformis – celtis – widespread, common – late spring/summer, leaves
Celticecis spiniformis – Celtis – widespread, common – late spring/summer, leaves
Celticecis supina – celtis, widespread, uncommon – late spring/summer, leaves
Clinodiplosis meibomiifoliae – tick trefoil - ? - Summer
Contarinia cerasiserotinae – black cherry - ? never personally observed – spring, stems
Contarinia cucumata – carya – rare? – late spring, leaves
Contarinia negundidis – box elder – uncommon, not sure if present in NE ohio – spring, petiole
Contarinia racemi – black cherry – uncommon – spring, flower raceme
Contarinia rumicis – European rumex sp – common? – late spring/early summer, seed
Contarinia verrucicola – Native Tilia – widespread, common – late spring/summer, leaves
Craneiobia tuba – cornus – rare – summer, leaves
Dasineura carbonaria – euthamia – rare? – summer, terminal
Dasineura collinsoniae – citronella horse balm – uncommon – early summer, leaves
Dasineura communis – sugar maple – scattered, locally common – late spring, leaves
Dasineura crataegibedeguar – hawthorn – uncommon, more common in NE Ohio – late spring, leaves
Dasineura gleditchiae – honey locust – common? – spring, leaves
Dasineura investita – wood nettle (Laportea) – widespread, locally common – very late spring/summer, leaves, stems, flowers
Dasineura involuta – wood nettle (Laportea) – rare – summer, terminal bud
Dasineura pellex – ash – widespread, common – spring, leaves
Dasineura pilosa – wood nettle (Laportea) – rare – summer, stems, leaves
Dasineura tumidosae – ash – rare (in my experience) – spring, leaves, petiole
Gliaspilota glutinosa – carya – widespread, uncommon – late spring/summer, leaf spot
Iteomyia salicisverruca – willows - ? not personally observed - ?
Lasioptera collinsonifolia – citronella horse balm - ? – late spring/summer, leaves
Macrodiplosis erubescens – oaks – uncommon? – spring, leaves
Macrodiplosis majalis – oaks – locally common – spring, leaves
Macrodiplosis nivepella – oaks – locally common - early spring, leaves
Macrodiplosis qoruca – oaks - ? – spring, leaves
Meunieriella – Smilax – uncommon – summer, leaves
Neolasioptera ambrosiae – giant ragweed – rare? – summer, stems
Neolasioptera asclepiae – Asclepias – rare? Never personally observed - ?, stems
Neolasioptera boehmeriae – Boehmeria – uncommon – summer,stems
Neolasioptera eupatorii – Eupatorium – uncommon? (not personally seen) – summer/fall, stems
Neolasioptera farinosa – Rubus – uncommon – summer, leaves
Neolasioptera impatientifolia – Impatiens – widespread, uncommon – summer, leaves-stems
Neolasioptera lycopi - Lycopus - ? never personally seen -?
Neolasioptera monardi – Monarda fistulosa - ?
Neolasioptera nodulosa – Rubus – uncommon?
Neolasioptera perfoliata – Eupatorium – uncommon – summer, stems
Neolasioptera verbesinae – Verbesina – widespread, common – summer, stems, leaf axils
Neolasioptera vernoniae – vernonia – widespread, common – summer, leaves, stems
Neolasioptera vitinea – grape – uncommon – summer, stems
Parallelodiplosis subtruncata – cornus – uncommon – summer, leaf spot
Peracecis fugitiva – celtis – rare SW Ohio – summer, leaves
Pilodiplosis helianthibula - sunflower - western ohio, common in prairie areas – early summer, leaves
Polystepha globosa – red oaks – widespread, common – summer, leaves
Polystepha pilulae – red oaks – widespread, common – very late spring-fall, leaves
Polystepha quercifolia – red oaks – uncommon – spring-fall, leaves
Rabdophaga ridigae – Salix – widespread, uncommon – fresh galls early summer, persistant throughout year, stems
Rabdophaga strobiloides – Salix - widespread, common – fresh galls early summer, persistent throughout year, terminal bud
Resseliella clavula – Cornus racemosa, other cornus - ? never personally observed – late spring, stems
Resseliella globosa – copperleaf – uncommon – early summer, stems
Resseliella liriodendri – Liriodendron – widespread, common – summer, fall, leaves
Resseliella tulipiferae – Liriodendron - summer, leaves
Rhopalomyia anthophila – Solidago – Rare – late summer, flowers
Rhopalomyia capitata – Solidago – common – summer, terminal bud
Rhopalomyia fusiformae – Euthamia – uncommon – summer, multiple plant parts
Rhopalomyia hirtipes - Solidago junacea - ? never personally seen, summer, terminal bud
Rhopalomyia pedicellata – Euthamia – uncommon – summer, multiple plant parts
Rhopalomyia solidaginis – Solidago – widespread, common – summer, terminal bud
Taxodiomyia cupressiananassa – Taxodium – more common as you move southwest, very common in Cincinnati – summer/fall, leaves
Schizomyia eupatoriflorae – Ageratina – widespread, uncommon – late summer, flowers
Schizomyia impatientis – Impatiens – widespread, uncommon – summer, flowers
Schizomyia racemicola – solidago – common – summer, flowers
Schizomyia verbesinae – Verbesina – rare? – summer, flowers
Vitisiella brevicauda – native grapes – uncommon – late spring/summer, stems, leaves, tendrils

Gall Mites
Acalitus ferrugineum – beech – widespread, common – late spring/summer, leaves
Acalitus longisetosus – River birch - ? never personally observed – spring/summer, leaves
Aceria campestricola – elm – common? – spring, leaves
Aceria caryae – carya – common – late spring/summer, leaves
Aceria caulis – black walnut – widespread, common – early summer, petiole
Aceria celtis – celtis – widespread, common – throughout year, witches broom
Aceria cephalanthi - buttonbush – uncommon – early summer, leaves
Aceria cinereae – butternut – common on host, but host is uncommon – early summer leaves
Aceria dina – black tupelo – widespread, uncommon – late spring/summer, leaves
Aceria elongata – sugar maple – rare? More common north and east – spring/summer, leaves
Aceria fraxini – ash – widespread, common – spring/summer - leaves
Aceria fraxiniflora – ash – uncommon – spring, flowers
Aceria major – red maple – common, but rarer than other maple erineum mites – spring/summer, leaves
Aceria modesta – sugar maple – common – spring/summer, leaves
Aceria mori – mulberry – uncommon – throughout year, witches broom
Aceria negundi – box elder – widespread, common – spring/summer, leaves
Aceria nyssae – black tupelo – widespread, common – spring/summer, leaves
Aceria parulmi – elm – widespread, common – spring/summer, leaves
Aceria parapopuli – Populus - ? never personally observed - ?
Aceria querci – bur oak - widespread, common – Late spring/summer, leaves
Aceria theospyri – Persimmon – common on persimmon but the tree is rare – late spring/summer, leaves
Aceria trinema – red oaks esp pin oak – widespread, common – most visible mid-spring, leaves
Aceria triplacis – white oak – widespread, common – late spring/summer, leaves
Aceria vaga – carya – uncommon? – late spring/summer - leaves
Aculops aenigma – Salix – uncommon? – late spring?/Summer?, Flower buds
Aculops euphorbicolus - Euphorbia corrolata - ?, probably more common in the Western part of the state where the host is more common
Aculops rhois – poison ivy - widespread, common – late spring/summer, leaves
Aculus minutissimus – red maple – rare, I have only seen in NE ohio – spring/summer, leaves
Aculus leionotus – river birch – uncommon – spring/summer, leaves
Aculus tetranothrix – Salix sp – widespread, common – late spring/summer, leaves
Cenalox nyssae – tupelo – uncommon? – late spring/summer, leaves
Eriophyes aceris – silver maple – uncommon – spring/summer, leaves
Eriophyes cerasicrumena – black cherry – widespread, common – early spring/summer, leaves
Eriophyes emarginatae – choke cherry and other native cherries – widespread, common – very early spring/summer, leaves
Eriophyes laevis – Alder – uncommon – spring/summer, leaves
Eriophyes leiosoma – European Tilia - ? – spring?, leaves
Eriophyes pyri – callery pear - ? – spring, leaves
Eriophyes tiliae – Native and European Tilia – widespread, common, spring/summer, leaves
Epitrimerus marginemtorquens – callery pear – common - spring/summer, leaves
Phyllocoptes didelphis – Populus – ? never personally observed - ?
Vasates aceriscrumena – sugar maple – widespread, common – early spring, summer, persisting into fall, leaves
Vasates quadripedes – red and sugar maple – widespread, common – early spring/summer, leaves
Hemipterans
Adelges abietis – firs – uncommon - ?, stems
Colopha ulmicola – elms – widespread, common – spring/early summer - leaves
Daktulosphaira vitifoliae – grape – widespread, common – very late spring/summer - leaves
Eriosoma americanum – elm – uncommon – spring/early summer - leaves
Hamamelistes spinosus – witch hazel, river birch – uncommon, more common on river birch – spring/summer - leaves
Hormaphis hamamelidis – witch hazel – widespread, common – spring/summer leaves
Kaltenbachiella ulmifusa - slippery elm -late spring, summer, leaves
Melaphis rhois – sumac – widespread, common – summer, leaves/petiole
Mordwilkoja vagabunda – Populus – rare? – spring, terminal bud
Pachypsylla celtidisinteneris – celtis - ? (not personally seen) - ?, stems
Pachypsylla celtidismama – celtis - widespread, common – late spring/fall, leaves
Pachypsylla celtidisumbilicus – celtis – Western Ohio, common – late spring/summer, leaves
Pachypsylla celtidisvesicula – celtis – widespread, common – late spring/summer, leaves
Pachypsylla venusta – Celtis – widespread, common – late spring/summer, petiole
Pemphigus populicaulis – Populus sp – common – summer, leaf base/petiole
Pemphigus populitransversus – Populus sp – uncommon – summer, petiole
Phylloxera caryaeavellana - uncommon, spring, leaves
Phylloxera caryaecaulis - carya - uncommon - spring, leaves and petioles
Phylloxera caryaeglobuli - carya - common - spring, leaves
Phylloxera caryaegummosa – Hickory – uncommon– spring, leaves
Phylloxera caryaemagna - ? - spring, leaves
Phylloxera caryaeren - carya - spring, leaves
Phylloxera caryaesemen - bitternut – common – very late spring/summer, leaves
Phylloxera caryaesepta - carya - uncommon– spring, leaves
Phylloxera flavoconica - carya - rare - spring, leaves
Phylloxera foveola - carya - rare, spring, leaves
Phylloxera killianae - carya - uncommon– spring, leaves
Phylloxera picta - carya - uncommon– spring, leaves
Phylloxera subelliptica – carya - ? – spring, leaves/petioles
Phylloxerina nyssa – tupelo – uncommon – late spring, leaves
Tetraneura nigriabdominalis – elms - ? - ?
Tetraneura ulmi – Elms – Widespread, common – late spring/summer, leaves

Fungi
Apiosporina morbosa – choke cherry – locally common - stems
Cronartium quercuum – pines? – not personally observed - stems
Diaporthe sp. – Bitternut – widespread, uncommon, but will cover entire trees where present – visible year round, stems
Gymnosporangium juniperi-virginianae – Junipers – Locally common – spring, stems/leaves
Taphrina caerulescens – red oaks – widespread, common – spring/summer, leaves
Taphrina communis – native plums - ? never personally observed – spring, fruits
Taphrina deformans – peach – uncommon – spring, leaves
Taphrina farlowii – black cherry – common – spring, leaves/fruits
Taphrina johansonii – Populus - ? not personally observed - ?, fruits
Taphrina populina – Populus – common – spring, leaves
Taphrina polystichi – Christmas fern – common – spring, leaves
Taphrina ulmi – elm – uncommon – spring, leaves

Other
Agrobacterium sp. – various hosts – bacteria, uncommon, often in cultivated situations – visible year round
Agromyza deserta – celtis – agromyzid fly - ?
Ecdytolopha insiticiana – black locust – moth, common? - ?
Ectoedemia populella – Populus – moth - ? – summer, petiole/leaf base
Eurosta solidaginis – Solidago – fly, widespread, common – summer, stem
Gnorimoschema gallaesolidaginis – solidago – moth, uncommon – summer, stem
Heliozela aesella – grape - ? (not personally observed) – early summer, leaves
Pontania sp – Salix sp - Sawfly - Leaves, summer
Urophora cardui – thistles – fly ? never personally observed

Undescribed gall wasps
Q alba bead gall – common – early summer/fall, leaves
Q muehlenbergii - white midrib gall – uncommon – summer, leaves
Q velutina - yellow midrib gall – uncommon – late summer, leaves
Q stellata - "phylloteras-like" spangle - rare, seen once in S Union county – late summer, leaves

Undescribed gall midges
Contarinia - box elder – uncommon – spring, leaves
Contarinia sp - oak - vein gall uncommon – spring, leaves
Polystepha? - red oaks - cone gall, uncommon – late summer, leaves

Uncertain presence in Ohio (my to-find list)
Wasps

Acraspis longicornus - post oak
Amphibolips coelebs – Red oaks
Amphibolips globulus - blackjack oak - unlikely given known distribution and rarity of host in Ohio
Amphibolips murata - blackjack oak - unlikely given known dist. and rare host
Amphibolips tinctoriae
Andricus clarkei - q alba q stellata
Andricus deciduuatus – q. bicolor
Andricus murtfeldtae - q stellata
Antistrophus jeanae - Silphium perfoliatum
Antistrophus laciniatus - Silphium lacinatum - probably unlikely, host rare in ohio
Antistrophus meganae - S. terebinthinaceum
Antistrophus minor - S. lacinatum - unlikely, host rare
Antistrophus rufus - S. lacinatum - unlikely, host rare
Antistrophus silphii – S.integrifolium/S. lacinatum - unlikely hosts rare
Aulacidea abdita - Latuca canadensis
Aulacidea annulata - Latuca/Prenanthes
Aulacidea harringtoni - Latuca floridana
Aulacidea nabali - Nabalus
Aulacidea pilosellae - Pilosella
Aulacidea podagrae - Latuca sp.
Aulacidea tumida - Latuca sp.
Biorhiza caepuliformis - red oaks
Callirhytis balanaspis - red oaks, esp black oak
Callirhytis balanoides - black oak
Callirhytis balanopsis - red oaks - probably unlikely
Callirhytis balanosa - red oaks
Callirhytis crypta - red oaks
Callirhytis electrea -chestnut oak
Callirhytis ellipsoida - swamp white oak
Callirhytis elliptica - q alba
Callirhytis excavata – Red oaks
Callirhytis florensis - blackjack oak - unlikely because of rarity of host
Callirhytis glomerosa - red oaks
Callirhytis hopkinsi - shingle oak
Callirhytis lapillula - swamp white oak
Callirhytis marginata - red oaks
Callirhytis medularis - red oaks
Callirhytis perrugosa - q alba
Callirhytis rubida - red oaks
Callirhytis rugulosa – red oaks
Callirhytis seminosa - red oaks
Callirhytis subcostata - post oak
Callirhytis tuberosa - shingle oak
Callirhytis tubicola - post oak
Callirhytis vaccinii - post oak
Diastrophus bassettii - Rubus procumbens
Diastrophus fragariae - Fragraria virginiana
Diastrophus niger - Potentilla canadensis
Diastrophus potentillae - Potentilla canadensis/ P. simplex
Diastrophus radicum - Rubus corchorifolius and R. occidentalis
Diastrophus turgidus – Rubus strigosus
Diastrophus niger – cinquefoil
Diplolepis fusiformans subsp. fusiformans - Rosa arkansana/Rosa blanda/Rosa woodsii - probaly unlikely hosts rare or not present in Ohio
Diplolepis gracilis – Rose - unlikely given known range
Diplolepis ignota – Rose, R. carolina most prevelant Ohio host
Diplolepis nodulosa – Rose - unlikely
Diplolepis pustulatoides - rose
Diplolepis radicum - R. carolina
Diplolepis spinosa – Rose - unlikely
Diplolepis tuberculosa - rose -
Disholcaspis bassetti - bur oak / swamp white oak
Disholcaspis spongiosa - post oak / white oak
Disholcaspis terrestris - post oak/white oak
Dryocosmus favus - red oaks
Eumayria enigma - red oaks
Eumayria floridana - red oaks
Eumayria saltata - red oaks
Holocynips badia - white oaks
Holocynips maxima - white oaks
Loxaulus beutenmuelleri - red oak
Loxaulus humilis - post oak
Loxaulus illinoisensis - white oak/bur oak
Loxaulus vaccinii - post oak
Melikaiella corrugis - red oaks
Melikaiella fructicola - red oaks
Melikaiella fructuosa - red oaks
Melikaiella ostensackeni
Melikaiella papula - red oaks
Neuroterus affinis - Q. prinoides
Neuroterus bassettii - q alba
Neuroterus consimilis - q alba
Neuroterus contortus - post oak
Neuroterus distortus - swamp white oak / chestnut oak
Neuroterus escharensis - q alba / q bicolor
Neuroterus exiguissimus - q alba
Neuroterus exiguus - post oak
Neuroterus fugiens – q alba
Neuroterus gillettei - post oak
Neuroterus niger - white oaks
Neuroterus pallidus - white oaks
Neuroterus papillosus - swamp white oak / chestnut oak
Neuroterus perminimus - q alba / q bicolor
Neuroterus rileyi - post oak /chestnut oak
Neuroterus tectus - chestnut oak
Neuroterus vernus - bur oak
Odontocynips nebulosa - post oak
Philonix gigas - post oak
Philonix pallipes - q alba
Phylloteras sigma – q alba
Sphaeroteras melleum - post oak
Trigonaspis polita - q alba / q stellata - unlikely given known geography
Trigonaspis quercusforticorne – bur oak
Trigonaspis radicola - q alba / q stellata
Zapatella davisae - red oaks
Zapatella oblata - scarlet oak
Zapatella quercusmedullae - red oaks
Zopheroteras cuneatum – q alba fall
Midges
Ametrodiplosis geminata – Mountain mint, midsummer
Asphondylia autumnalis – Helenium –
Asphondylia azaleae – Rhododendron –
Asphondylia diervillae – Northern Bush Honeysuckle –
Asphondylia hydrangeae – Hydrangea arborescens
Asphondylia lacinariae – Liatris pycnostachya –
Asphondylia ratibidae - Ratibida
Asphondylia recondita – Asters, rosette gall
Asphondylia rosulata - solidago
Asphondylia sambuci – Elderberry bud gall
Asphondylia smilacinae – Smilacina racemosa
Asphondylia silva – Bluestem goldenrod –
Asphondylia thalictri – Thalictrum sp.
Caryomyia albiposa – Carya - spring
Caryomyia arcuata – Carya and rest of Caryomyia below
Caryomyia caminata –
Caryomyia cynipsea –
Caryomyia glebosa
Caryomyia hirtiglobus
Caryomyia ovalis
Caryomyia spherica
Caryomyia striolacrustum
Caryomyia turbanella
Caryomyia turbinata
Cecidomyia squamulicola – hazel catkins
Celticecis wellsi - celtis
Clinodiplosis apocyni – hemp dogbane
Clinodiplosis hastate – Verbena hastata
Clinodiplosis lappa – Spiraea salicifolia –
Clinodiplosis rhododendri – Rhododendron
Clinodiplosis spiraeina – spiraea alba
Contarinia citrina – Tilia americana
Contarinia clarkei – Spiraea alba
Contarinia juniperiana – Juniper sp.
Contarinia nucicola – hickory –
Contarinia pyrivora – Pyrus communis
Contarinia virginianiae – chokecherry
Cystiphora canadensis – Nabalus
Dasineura americana – Galium asprellum
Dasineura anemone – Anemone canadensis
Dasineura aromaticae – Mentha arvensis, Mentha spicata
Dasineura dentatae – chestnut
Dasineura folliculi – Solidago
Dasineura johnsoni – Vitis spp
Dasineura laquerrarum – Mountain mint
Dasineura lepidii – Lepidium (peppergrass)
Dasineura lysimachiae – Lysimachia quadrifolia, terrestris
Dasineura mali – Apple
Dasineura parthenocissi – Virginia Creeper
Dasineura piperitae – Mentha gentilis, Mentha piperita
Dasineura plicata – Salix sp.
Dasineura porrecta – Ulmus americana
Dasineura pseudacaciae – black locust
Dasineura pudibunda - hornbeam
Dasineura pyri – Pyrus communis
Dasineura rhodophaga – Roses (cultivated)
Dasineura sassafras – Sassafras
Dasineura semenivora – Viola spp
Dasineura serrulatae – Alder bud gall
Dasineura smilacifolia – Smilax spp
Dasineura toweri – Hypericum mutilum
Dasineura trifolii – Clover
Dasineura ulmae – Ulmus americana
Dasineura urnicola – Urtica dioica
Harmandia/Harmandiola - populus
Iteomyia salicifolia – Salix sp –
Janetiella asplenifolia – Myrica asplenifolia
Janetiella ulmii - Elm
Lasioptera cylindrigallae – Euthamina
Lasioptera excavata – Crataegus sp.
Lasioptera humulicaulis – Humulus
Lasioptera lactucae – Lactuca Canadensis
Lasioptera lorrainae – Mountain mint
Lasioptera spiraeafolia – Spiraea densiflora, douglasii, salicifolia
Macrodiplosis castaneae – Chestnut
Meunieriella aquilonia – honey locust
Neolasioptera brevis – honey locust
Neolasioptera clematidis – Clematis
Neolasioptera convolvuli – Bindweed
Neolasioptera cornicola – Cornus stononifera
Neolasioptera desmodii – tick trefoil
Neolasioptera erigeroni - fleabane
Neolasioptera fontagrensis – Celastrus scandens
Neolasioptera galeosidis – Galeopsis
Neolasioptera hibisci – Hibiscus
Neolasioptera linderae – Lindera benzoin
Neolasioptera menthae – Mentha sp
Neolasioptera nodulosa – native Rubus
Neolasioptera pierrei – Elderberrry
Neolasioptera rudbeckiae - Rudbeckia
Neolasioptera triadenii – Triadenum virginicum
Neolasioptera viburnicola – viburnum dentatum
Olpodiplosis helianthi – Helianthus
Obolodiplosis robiniae – black locust
Polystepha podagrae - black oak
Polystepha pustulata black oak
Polystepha pustuloides - black oak
Polystepha serrata - scarlet oak
Polystepha simpla - black oak
Polystepha sobrina - black oak
Prodiplosis myricae – Myrica cerifera
Prodiplosis morrisi – Populus sp.
Prodiplosis platani – Platanus occidentalis
Prodiplosis violicola – Viola spp
Rabdophaga cephalanthi – Cephalanthus occidentalis
Rabdophaga salicisbatatus - Salix
Rabdophaga saliciscornu – Salix humilis
Rabdophaga saliciscoryloides - Salix
Rabdophaga salicistriticoides – Salix cordata, Salix humilis -
Rabdophaga salicisrhodoides – Salix sp
Rhopalomyia castanae – Chestnut
Rhopalomyia chrysanthemi –cultivated chrysanthemums
Rhopalomyia clarkei – Solidago rugosa, S. altissima
Rhopalomyia gemmaria - asters
Rhopalomyia grossulariae – Ribes grossularia
Rhopalomyia inquisitor – Solidago
Rhopalomyia lobata – Euthamia
Rhopalomyia hirtipes – Solidago junacea
Rhopalomyia sp. – Asters, floret gall
Rhopalomyia strobiligemma – Panicled aster
Rhopalomyia weldi – Bigleaf Aster
Sackenomyia commota – virburnum
Sackenomyia viburnifolia – viburnum dentatum
Schizomyia umbellicola – Elderberry
Schizomyia viburni – Viburnum
Mites
Acalitus brevitarsus - Alder
Acalitus calycophthirus - paper birch - unlikely given rarity of host
Acalitus phyllereus – Alder
Acalitus rudis – Paper birch
Aceria kuko - Lycium
Aceria spicati - Mountain maple
Aculus craspedobius –willows
Aculops ailanthii – Ailanthus
Cenalox conyssae - tupelo
Colomerus vitis – Grape
Eriophyes betulae – birch buds
Eriophyes helicantyx - bracken fern
Eriophyes paraviburni – viburnum
Eriophyes rhoinus - sumac
Phytopus avellana – hazel
Hemiptera
Pachypsylla celtidisgemma – celtis stems
Pachypsylla celtidisinteneris – celtis stems
Phylloxera caryaefallax - carya - leaf
Phylloxera caryaefoliae - carya - leaf
Phylloxera caryaescissa - carya - leaf
Phylloxera caryaevenae - carya - leaf
Phylloxera conica - Carya - leaf
Phylloxera crypta - carya - leaf
Phylloxera devastatrix - pecan - leaf
Phylloxera falsostium - carya - leaf
Phylloxera myristica - carya - leaf
Phylloxera notabilis - carya - leaf
Phylloxera paludis
Phylloxera pilosula - carya - leaf
Phylloxera russellae - carya - leaf
Phylloxera spinosa - carya - leaf
Phylloxera stoetzelae - carya - leaf
Phylloxera subelliptica - carya - petiole
Phylloxera symmetrica - carya - leaf
Phylloxera wiedenmanni - carya - leaf
Phylloxera williamsi - carya - leaf
Fungi
Taphrina americana - birch
Taphrina betulina - birch
Taphrina carveri – maple
Taphrina coryli - hazel
Taphrina ostryae - Hophornbeam
Taphrina robinsoniana – alder
Taphrina wisneri - cherry
Pear rust
Other
Euura proxima - Salix - sawflies
Euura exiguae - Salix - sawflies
Lixus musculus – Polygonum - beetle
Pseudomonas savastanoi – Forsythia - bacteria

Publicado em 29 de junho de 2021, 11:41 PM por calconey calconey | 9 comentários | Deixar um comentário

18 de junho de 2021

Midge Gall Wishlist

A list of midge galls I would like to find, some of which are present here to some extent, but none widely reported.

Presented here in case anybody else wants to look for some of these. List relevant for Eastern/Midwest US.

Ametrodiplosis geminata – Mountain mint, midsummer - bud gall
Asphondylia autumnalis – Helenium autumnale - swollen, globular leafy bud gall
Asphondylia azaleae – Rhododendron – Enlarged bud
Asphondylia diervillae – Northern Bush Honeysuckle – Enlarged deformed bud or fruit (often sickle-shaped)
Asphondylia hydrangeae – Hydrangea arborescens – enlarged bud
Asphondylia lacinariae – Liatris pycnostachya – spherical bud gall
Asphondylia recondita – Asters - rosette gall
Asphondylia sambuci – Elderberry - bud gall
Asphondylia smilacinae – Smilacina racemosa – deformed berry
Asphondylia silva – Bluestem goldenrod –
Asphondylia thalictri – Thalictrum sp. – enlarged seed pod
Celticecis wellsi - Celtis occidentalis - leaf gall
Clinodiplosis apocyni – hemp dogbane - swollen flower bud
Clinodiplosis lappa – Spiraea salicifolia – narrow elongate clustered buds
Clinodiplosis meibomiifoliae – tick trefoil – swollen, aborted bud gall
Clinodiplosis rhododendri – Rhododendron – rolled distorted young leaf
Clinodiplosis hastata – Verbena hastata – marginal leaf roll
Contarinia citrina – Tilia americana – swollen young bud or twig
Contarinia juniperiana – Juniper sp. – Slight swelling at base of needle
Contarinia nucicola – hickory – surface swelling of husk
Contarinia pyrivora – Pyrus communis – enlarged sterile fruitlet
Cystiphora canadensis – Nabalus – circular leaf spot
Dasineura americana – Galium asprellum – swollen aborted bud
Dasineura anemone – Anemone canadensis – slightly englarged bud
Dasineura aromaticae – Mentha arvensis, Mentha spicata – crumpled, folded, young leaf
Dasineura collinsoniae – Collinsonia canadensis – onion shaped leaf gall
Dasineura dentatae – chestnut – flat, circular, warty growth, usually between veins
Dasineura folliculi – Solidago
Dasineura johnsoni – Vitis spp – deformed fruit
Dasineura laquerrarum – Mountain mint – small bud gall
Dasineura lepidii – Lepidium (peppergrass) infested swollen seed capsule (darker color)
Dasineura lysimachiae – Lysimachia quadrifolia, terrestris – swollen terminal bud
Dasineura mali – Apple – Curled leaf
Dasineura meibomiae – tick trefoil – hard, elongate-oval bud gall
Dasineura parthenocissi – Virginia Creeper - fleshy vein fold gall
Dasineura piperitae – Mentha gentilis, Mentha piperita – Swollen bud
Dasineura plicata – Salix sp. – Rolled leaf
Dasineura porrecta – Ulmus americana – deformed samara
Dasineura pseudacaciae – black locust – tightly folded swollen leaflet
Dasineura pyri – Pyrus communis – curled leaf
Dasineura rhodophaga – Roses (cultivated) – curled and stunted buds
Dasineura sassafras – Sassafras – curled leaf/leaf margin
Dasineura semenivora – Viola spp – enlarged, deformed, onion shaped fruit
Dasineura serrulatae – Alder bud gall
Dasineura smilacifolia – Smilax spp – reddened marginal leaf roll
Dasineura toweri – Hypericum mutilum – enlarged seed capsule
Dasineura trifolii – Clover – folded and swollen along midvein, on introduced European Trifolium species
Dasineura ulmae – Ulmus americana – enlarged leaf bud
Dasineura urnicola – Urtica dioica - Pale green midrib or vein gall, subsessile, diameter yi inch, midge
Iteomyia salicifolia – Salix sp – spherical gall protruding from only one side of leaf
Janetiella asplenifolia – Myrica asplenifolia – swollen vein, severe leaf distortion
Lasioptera collinsonifolia – Collinsonia – swollen midrib/lateral vein
Lasioptera cylindrigallae – Euthamina – tapered stem swelling (base of stem)
Lasioptera excavata – Crataegus sp. – Circular leaf spot
Lasioptera humulicaulis – Humulus - swollen stem
Lasioptera lactucae – Lactuca canadensis - tapered stem swelling, woody
Lasioptera lorrainae – Mountian mint - spherical stem gall
Lasioptera spiraeafolia – Spiraea densiflora, douglasii, salicifolia – circular leaf spot
Macrodiplosis castaneae – Chestnut – smooth, globular leaf fold
Meunieriella aquilonia – honey locust – ovoid leaf blister
Neolasioptera brevis – honey locust – swollen stem
Neolasioptera clematidis – Clematis – tapered stem swelling
Neolasioptera convolvuli – Bindweed – Tapered stem swelling
Neolasioptera cornicola – Cornus stolonifera – tapered ridged stem swelling
Neolasioptera desmodii – tick trefoil – swollen stem
Neolasioptera fontagrensis – Celastrus scandens – swollen stem
Neolasioptera galeosidis – Galeopsis – irregular stem swelling
Neolasioptera hibisci – Hibiscus – swollen stem
Neolasioptera linderae – Lindera benzoin – irregular stem swellling
Neolasioptera lycopi – Lycopus – swollen stem
Neolasioptera menthae – Mentha sp. – Swollen stem
Neolasioptera monardi – Monarda fistulosa – tapered stem swelling
Neolasioptera nodulosa – native Rubus – swollen stem
Neolasioptera pierrei – Elderberrry - stem swelling
Neolasioptera triadenii – Triadenum virginicum – irregular stem swelling
Neolasioptera viburnicola – viburnum dentatum – swollen stem
Olpodiplosis helianthi – Helianthus - tubular node gall
Prodiplosis myricae – Myrica cerifera – swollen bud, rolled leaf
Prodiplosis morrisi – Populus sp. – rolled leaf/curled leaf margin
Prodiplosis platani – Platanus occidentalis – rolled leaf margin/distorted leaf
Prodiplosis violicola – Viola spp. – rolled deforemed leaf
Resseliella tulipiferae – Tulip tree – leaf vein swelling
Rabdophaga cephalanthi – Cephalanthus occidentalis – swollen stem
Rabdophaga saliciscornu – Salix humilis (?) – lateral bud deformed into thin-walled horn-like structure
Rabdophaga salicistriticoides – Salix cordata, Salix humilis - forshortened twig, buds close together, resembling a wheat head
Rhopalomyia castanae – Chestnut – enlarged clustered buds
Rhopalomyia chrysanthemi – cone shaped leaf galls cultivated chrysanthemums
Rhopalomyia grossulariae – Ribes grossularia – enlarged deforemed bud
Rhopalomyia lobata – Euthamia - spongy apical gall
Rhopalomyia hirtipes – Solidago junacea - hard, globular apical gall
Rhopalomyia strobiligemma – Panicled aster – rosette gall
Rhopalomyia weldi – Bigleaf Aster, Spring
Sackenomyia commota – virburnum circular leaf blister
Sackenomyia viburnifolia – viburnum dentatum – purple vein swelling
Schizomyia umbellicola – Elderberry large unopened flower
Schizomyia viburni – Viburnum enlarged blossom or bud

Publicado em 18 de junho de 2021, 06:59 PM por calconey calconey | 2 comentários | Deixar um comentário

10 de março de 2021

Tips for Gall Hunting

Tips on Gall Hunting; Where and When to Hunt for Galls

Think about sites that have mature trees with low-hanging branches. Sites that have been quite good for me include city parks, cemeteries, and forests with lots of edges where the forest meets open ground, such as fields or prairies. Galls are often on large tree species, so closed forest environments can be more difficult, especially if you are looking for galls on shade-intolerant species such as oaks. Forests with edges next to prairies are particularly fantastic as you can find galls on the trees and the herbs.

Areas of high human impact have some influence on the frequency of gall species, at least the more uncommon species. However, do not disregard manicured areas such as arboretums, city parks, golf courses or even tree lawns as they will often have galling species. For example, I found 20 or so gall species on the highly managed "Oval" on Ohio State University's main campus. Diversity does tend to be lower in these spots but do not discount them as a place to look.

If you just want to find a gall (any gall), seek out sites with plant species that have a huge diversity of galls. Examples of species with large gall diversity include; Quercus (particularly alba, coccinea, stellata), Carya (in late summer many of these will have a half dozen gall species), Celtis (many here in Ohio have 5-6 species on a single tree, although I am not sure how far east some of these go), and Solidago/Euthamia. iNat distribution maps and geotags can be useful for finding these sites. Another recommendation if you are looking for a particular host is to do a search for the common name of the host on your state's DNR website, they will often list some of the common species that are located at a given nature preserve or state park.

If you are looking for a specific gall, try to find a site where the host is regionally native and common. Trees that are native and more common in a region will tend to have higher diversity. For example, Quercus macrocarpa and Quercus muehlenbergii are native but uncommon in NE Ohio. I have found far fewer gall species on them in this area compared to western Ohio where both species are much more common. Hackberry, Celtis occidentalis, is another example of this, it becomes much more common as you move south and west in Ohio and the diversity of galls seems to increase in that direction.

If you notice that a particular tree has a large number of galls, keep visiting that tree repeatedly through the course of the year and even over multiple years, and you will likely run into new species. It seems like some trees have something like a susceptibility factor or alternatively, some kind of environmental niche like wind direction that seems to make them more galled than others. These trees have been a rich source of new species for me.

Mid to late summer (July-September) is the best season for hunting galls in Ohio. The highest number of species will be around during that time. However, if you want to find rarer or underreported stuff, look for galls during what might seem like the offseason. For example, early spring (March-May) has many interesting oak bud galls. October has another subset of oak galls that appear just as the leaves are turning that don't often get reported. November and December after the leaves fall can be a good time to look for galls in the leaf litter of more closed forests.

Publicado em 10 de março de 2021, 06:30 PM por calconey calconey | 8 comentários | Deixar um comentário

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