Arquivos de periódicos de junho 2022

14 de junho de 2022

Systema Dipterorum's Checklist of Limnophora (Diptera: Muscidae) north of Mexico

The species of Limnophora and related genera are poorly known and a nomenclatural mess. The Systema Dipterorum probably serves as the only modern or recent taxonomic authority for such genera, at least for the Nearctic region. The work in Stone et al. (1965) hardly differs from this catalogue except for the addition of the species L. invada from California described by Huckett in 1966. Nearctica.com includes L. corvina even though it is not known to occur north of Mexico, while Systema Dipterorum lists it as neotropical. Of course, it is always possible that it exists here but is unrecorded.

Checklist (10-11 species):
[Limnophora corvina (Giglio-Tos, 1893: 7): Mexico, Puerto Rico, Peru, ?Brazil. Chile]
Limnophora discreta Stein, 1898: 204 - Alaska to California and New Mexico, British Columbia to Newfoundland, s. to Ill.andFla.
Limnophora incrassata Malloch, 1919: 299 - Alaska,Washington, Montana, Oregon, Wyo.,Colorado
Limnophora invada Huckett, 1966: 285 - California
Limnophora femorata (Malloch, 1913: 603) - Arizona, New Mexico, BajaCalif.
= Tetramerinx femorata Malloch, 1913: 603
Limnophora narona (Walker, 1849: 945) - Washington to Alberta & Quebec, s. to Chile & Argentina; Bermuda, West Indies
= Anthomyia narona Walker, 1849: 945
= Anthomyia prominula Thomson, 1869: 550
= Homalomyia dentata Bigot, 1885: 284
= Limnophora cyrtoneurina Stein, 1898: 203
= Limnophora indecisa Stein, 1901: 198 (unavailable)
= Limnophora transmutans Stein, 1901: 214 (unavailable)
Limnophora nigripes (Robineau-Desvoidy, 1830) - Alaska to Greenland, s. to California,Quebec, and Newfoundland, also S.Dak.
= Limosia nigripes Robineau-Desvoidy, 1830: 541
Limnophora groenlandica Malloch, 1920: 147 - West Greenland
Limnophora rotundata (Collin, 1930: 256) - w Greenland
= Pseudolimnophora rotundata Collin, 1930: 256
Limnophora sinuata Collin, 1930: 256 - Newfoundland, Greenland, Iceland,Favoes,Norway
= Limnophora islandica Lyneborg, 1965: 215
Limnophora uniseta Stein, 1916: 94 - Alaska to Greenland,also British Columbia, Washington, n.Europe
= Limnophora quadripunctata Curtis, 1837: 264 (nomen nudum)
= Limnophora coenosiaeformis Schnabl, 1889: 336 (nomen nudum)
= Limnophora nilotica Schnabl, 1889: 335 (nomen nudum)
= Limnophora parallelinervis Schnabl, 1889: 335 (nomen nudum)
= Limnophora lunulata Santos Abreu, 1976: 97
= Limnophora tenuiornata Santos Abreu, 1976: 91

Reference
Bigot, J.M.F. 1885. Diptères nouveaux ou peu connus. 25 partie, XXXIII: Anthomyzides nouvelles. Annales de la Société Entomologique de France (1884) (6) 4: 263-304.
Collin, J.E. 1930. A revision of the Greenland species of the anthomyid genus Limnophora sens. lat. (Diptera), with figures of the male genitalia of these and many other Palaearctic species. Transactions of the Royal Entomological Society of London 78: 255-281, 13 pls.
Curtis, J. 1837. A guide to an arrangement of British Insects: being a catalogue of all the named species hitherto discovered in Great Britain and Ireland. Second Edition, Greatly enlarged. J. Pigot, Sherwood and Co. & Marshall, London. vi pp. + 294 columns.
Huckett, H.C. 1932. The North American species of the genus Limnophora Robineau-Desvoidy, with descriptions of new species (Muscidae, Diptera). Journal of the New York Entomological Society 40: 25-76, 105-158, 279-338, 7 pls.
Huckett, H.C. 1966. New species of Anthomyiidae and Muscidae from California (Diptera).Proceedings of the California Academy of Sciences 34: 235-305.
Lyneborg, L. 1965. On Muscidae and Anthomyiidae (Diptera) from Iceland. With the description of two new species. Opuscula Entomologica 30: 211-226.
Malloch, J.R. 1913. Three new species of Anthomyiidae (Diptera) in the United States National Museum collection. Proceedings of the United States National Museum 45[2004]: 603-607.
Malloch, J.R. 1919. New species of flies (Diptera) from California. Proceedings of the California Academy of Sciences (4) 9: 297-312.
Malloch, J.R. 1920. Descriptions of new North American Anthomyiidae (Diptera). Transactions of the American Entomological Society 46: 133-196, 3 pls.
Santos Abreu, E. 1976. Monografia de los Anthomyidos de las Islas Canarias (Dipteros). Servicio del Aula de Cultura Elias Santos Abreu. 175 pp., 24 col. figs.
Schnabl, J. 1889. Contributions a la faune dipterologique. Trudy Russk. Ent. Obshch. 23: 313-347.
Stein, P. 1898. Nordamerikanische Anthomyiden. Beitrag zur Dipteren-fauna der Vereinigten Staaten. Berliner Entomologische Zeitschrift (1897) 42: 161-288.
Stein, P. 1901. Die Walker'schen aussereuropäische n Anthomyiden in der Sammlung des British Museum zu London. Zeitschrift für Systematische Hymenopterologie und Dipterologie 1: 185-221.
Stein, P. 1916. Die Anthomyiden Europas. Tabellen zur Bestimmung derGattungen und aller mir bekannten Arten, nebst mehr oder wenigerausfuhrlichen Beschreibungen. Archiv für Naturgeschichte (1915) (A) 81(10): 1-224.
Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, J.R. 1965. A catalog of the Diptera of America north of Mexico. Agric. Handbk 276, 1696 pp.
Thomson, C.G. 1869. Diptera. Species nova descripsit, pp. 443-614. In: Kongliga svenska fregatten Eugenies resa omkring jorden under befäl af C.A. Virgin, åren 1851-1853. 2 (Zoologi) 1, Insecta. "1868". P.A. Norstedt & Soner, Stockholm. 617 pp., pl. 9.
Walker, F. 1849. List of the specimens of dipterous insects in the collection of the British Museum. Part IV. British Museum (Natural History), London. Pp. [3] + 689-1172.

Posted on 14 de junho de 2022, 02:06 PM by aispinsects aispinsects | 6 comentários | Deixar um comentário

20 de junho de 2022

Systema Dipterorum's Checklist of Phaonia and Dolichophaonia (Diptera: Muscidae) north of Mexico

The "Phaoniini" of muscid flies in North America include some of the most diverse genera of the family in our area. Unfortunately, they are also very poorly known, often confused in nomenclature, and difficult to recognize from photographs. No checklist of the genus Phaonia north of Mexico has been made since Stone et al.'s catalogue, and Dolichophaonia was described in 1993, after which some Phaonia species in our area were almost silently transferred to. I have here compiled a list of species occuring in this area according to Systema Dipterorum, which keeps tabs on these taxa watchfully, including works published this year. It's one of the only options of sources we have to help make accessible checklists like these. Carvalho et al. made a catalogue of Muscidae of the Neotropical region in 2005. Next I will try to do this for Helina, which I also have specimens of, and might be an even more heterogenous genus than Phaonia.

Synopsis of Genera:
Genus Phaonia Robineau-Desvoidy, 1830: 482 (79 species)
In my trip to Massachusetts, I caught some flies and one of them keyed to Phaonia apicata (ssp. apicata? No nominal subspecies has been proposed) in Malloch (1923). That alone doesn't mean it is apicata; species not included in that key also should be accounted for, hence why I made this updated checklist, as notes for myself, but I would also like to share this with other people (I'm just not sure if I should be sharing this information like crazy, mostly due to copyright and stuff).
One of the defining characters for the genus Phaonia is the presence of hind tibia bare of setae except for a single well developed posterodorsal seta on the apical fourth or so. Other genera may also have this character, though, such as Hydrotaea (can we come to a consensus on the Hydrotaea-Ophyra issue please?!?!), Lophosceles, Muscina, and more. Helina almost always have a different configuration of the hind tibial chaetotaxy as Phaonia, with either more than 1 seta on the posterodorsal surface or with a single seta reduced in size or displaced. Arguably what's actually probably the more unique character of Phaonia is that many, maybe even most Phaonia also have 3 post-sutural dorsocentral setae, an exception among Muscidae, which usually have 4. Generally only Coenosiini and maybe some Helina have this character state. This can make them more difficult to separate from Anthomyiidae, however. In these cases, acrostichals also tend to be entirely lacking except for a prescutellar pair, which almost never occurs in Anthomyiidae. Most anthomyiids also have more than one posterodorsal seta on the hind tibiae. The crossveins on the wings may also be clouded with dark in Phaonia. The eyes can also be hairy.
Several additions to the fauna have been made since the keys from Malloch (1923) and Huckett (1965), and few synonymies. Ringdahl (1933) described P. bidentata. Hall (1937) described P. pudoa. Huckett (1965) of course described a few species. After Stone et al.'s catalogue, Huckett (1966) described several species from western North America. Huckett (1973) described P. cauta from Maine. Dialyta rufitibia, used in Huckett (1965), was synonymized with Phaonia flavitibia, and is consequently now Phaonia flavitibia. Phaonia morio Zetterstedt, used in Huckett (1965), was synonymized with P. lugubris. P. pruinosa became a junior primary homonym of Polietina concinna.
Not as straightforwardly, Phaonia pratensis has a markedly confused history of nomenclature. Most of it was summarized by Collin (1951) and Pont (1984). I haven't found it or its synonyms at all in Malloch (1923), Stone et al. (1965), or Huckett (1965), but it's listed to occur here by Systema Dipterorum. It turns out this species had been referred to as over 4 different species-group names. Collin (1951) described P. laetabilis, an additional species found among an assemblage also containing P. laeta Fallén in the UK. Later on, the valid name of P. laetabilis became P. trigonalis and that of P. laeta Fallén became P. pratensis. The lectotype of P. laeta turned out to be P. trigonalis itself though, and so Pont (1984), whom also explains all this, synonymized P. laetabilis with P. laeta. This should also imply that all of these names are just P. pratensis, since P. laeta is already a synonym of P. pratensis, but this was not made too clear. Systema Dipterorum lists both P. laeta and P. pratensis as valid names, but the ranges are printed exactly the same, suggesting that they meant for the two species to be the same. Therefore, I treat them all as P. pratensis here. I won't get into the other species-group names, but they are also synonyms. Collin (1951) denotes its morphology.

Genus Dolichophaonia Carvalho, 1993: 19 (2 species)
de Carvalho (1993) described the genus Dolichophaonia in their publication written in Portuguese. All the species assigned to the genus at the time were Neotropical, none occuring north of Mexico (yet?). Later on, our Phaonia texensis was provisionally transferred to the genus. Vockeroth (1996) shortly in his key to genera synonymized Dolichophaonia with Phaonia, suggesting there are no ways to separate females of that genus from Phaonia. This interpretation is reflected in the Muscidae portion in the Manual of Central America, written by Savage and Vockeroth (2015), which still considered Dolichophaonia a part of Phaonia. Couri & de Carvalho rejected this synonymy in 2002 during their key and taxonomic work of the Muscidae of the Neotropical region (that's also extremely difficult to access; I do not have it). It remains controversial if Dolichophaonia should remain a distinct genus. Systema Dipterorum considers it so. Fortunately, Malloch (1923) includes our Dolichophaonia species as Phaonia species and they are quite easily separated. Those species form a group in the key, suggesting a distinct relationship. At this time, only Dolichophaonia texensis appears to have been collected north of Mexico. Eventually, D. limbinervis will probably be found here. Systema Dipterorum broadly lists North America as part of its range anyway.

Checklist:
Genus Phaonia Robineau-Desvoidy, 1830: 482 (79 species)
Phaonia aberrans Malloch, 1919: 208 - New Jersey, Tennessee,North Carolina References: Malloch (1923)
Phaonia albocalyptrata Malloch, 1920: 267 - s.Alaska References: Malloch (1923), Huckett (1965)
Phaonia alpicola (Zetterstedt, 1845: 1401) - Alaska, Nunavut, Manitoba, Quebec,Europe,Palaearctic References: Huckett (1965)
= Aricia alpicola Boheman, 1844: 104 (nomen nudum)
= Aricia alpicola Zetterstedt, 1845: 1401
Phaonia alticola Malloch, 1923: 260 - Alaska, Nunavut,Yukon Territory References: Huckett (1965)
Phaonia antennalis Huckett, 1966: 296 - California
Phaonia apicalis Stein, 1914: 46 - Alaska,Michigan,n.Europe References: Huckett (1965)
Phaonia apicata Johannsen, 1916: 396 - Alberta, Iowa, Wisconsin,Michigan, Quebec, Labrador toGa. References: Malloch (1923)
+ssp. solitaria Stein, 1920: 15 - Wisconsin, New York
Phaonia atlanis Malloch, 1923: 279 - Quebec, Mich.to New York, s. to Miss.andFla.
Phaonia atrocitrea Malloch, 1923: 262 - Yukon Territory,Quebec References: Huckett (1965)
Phaonia atrocyanea Ringdahl, 1916: 234 - Alaska, Yukon Territory,Nunavut, Quebec, Labrador,Sweden References: Malloch (1923) (as citreibasis), Huckett (1965)
= Phaonia citreibasis Malloch, 1920: 268
Phaonia aurea Malloch, 1923: 256 - Washington
Phaonia azygos Malloch, 1923: 261 - New York
Phaonia basiseta Malloch, 1920: 133 - Alberta to Nunavut,s.to Alberta, South Dakota and Minnesota, e.Asia References: Malloch (1923), Huckett (1965)
Phaonia bidentata Ringdahl, 1933: 17 - Nunavut, Quebec,Labrador References: Huckett (1965)
Phaonia brevispina Malloch, 1923: 269 - Washington, to California,Idaho, Illinois to New Hampshire andVa.
Phaonia bysia (Walker, 1849: 936) - Alaska to Oreg., Alberta andIdaho, Michigan to Quebec, s. toGa. References: Malloch (1923), Huckett (1965)
= Anthomyia bysia Walker, 1849: 936
= Phaonia pallicornis Stein, 1920: 12
Phaonia caerulescens (Stein, 1898: 187) - Washington to Alberta, s.to California andUtah References: Malloch (1923)
= Aricia caerulescens Stein, 1898: 187
Phaonia californiensis (Malloch, 1923: 236) - California,Arizona
= Bigotomyia californiensis Malloch, 1923: 236
Phaonia cauta Huckett, 1973: 232 - Maine
Phaonia consobrina (Zetterstedt, 1838: 665) - Alaska to Greenland, s. to Quebec, Europe, n. Russia References: Malloch (1923), Huckett (1965)
= Anthomyza consobrina Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza consobrina Zetterstedt, 1838: 665
= Anthomyza vicina Zetterstedt, 1838: 673
= Aricia marmorata Zetterstedt, 1860: 6197
= Anthomyia serva Meigen of Lundbeck, 1898: 309
Phaonia coriatlanis Huckett, 1966: 297 - California
Phaonia curvinervis Malloch, 1923: 275 - N.S.
Phaonia curvipes (Stein, 1920: 19) - Michigan to Quebec, s.toN.C. References: Huckett (1965)
= Trichopticus curvipes Stein, 1920: 19
Phaonia deleta (Stein, 1898: 178) - Nunavut,California, Texas, Iowa to Quebec, s. toPa. References: Malloch (1923), Huckett (1965)
= Aricia deleta Stein, 1898: 178
Phaonia diruta (Stein, 1898: 188) - South Dakota,Kansas,Michigan to Connecticut, s. toGa. References: Malloch (1923)
= Spilogaster diruta Stein, 1898: 188
Phaonia dissimilis Malloch, 1923: 263 - Alaska References: Huckett (1965)
Phaonia errans (Meigen, 1826: 112) - Alaska to Newfoundland, s. toCalif. and New Jersey,Palaearctic References: Malloch (1923), Huckett (1965)
= Musca erratica Fallén, 1825: 77
= Anthomyia errans Meigen, 1826: 112
= Trennia nigricornis Robineau-Desvoidy, 1830: 484
= Anthomyza zetterstedt Bonsdorff, 1866: 273
= Yetodesia tinctipennis Rondani, 1866: 104
= Yetodesia manicata Rondani, 1871: 322
= Aricia girschneri Schnabl, 1888: 401
= Phaonia biseta Ringdahl, 1935: 31
+ssp. completa Malloch, 1923: 258 - New Hampshire
= Phaonia errans var. completa Malloch, 1923: 258
+ssp. luteva Walker, 1849: 934 - Nova Scotia
= Hyetodesia varipes Coquillett, 1900: 441
Phaonia fausta Huckett, 1965: 313 - Nunavut
Phaonia flava Stein, 1920: 6 - British Columbia, Oregon,California References: Malloch (1923)
Phaonia flavibasis Malloch, 1919: 208 - New York,New Hampshire References: Malloch (1923)
Phaonia flavitibia (Johannsen, 1916: 394) - Wisconsin to Quebec,s. toN.C. References: Huckett (1965) (as Dialyta)
=Dialyta flavitibia Johannsen, 1916: 394
=Dialyta rufitibia Stein, 1920: 23
Phaonia fraterna Malloch, 1923: 251 - New Hampshire
Phaonia fuscana Huckett, 1965: 906 - Washington, Oregon, Wisconsin to Quebec, s. to Tex.andGa.
= Spilogaster fusca Stein, 1898: 189
Phaonia fuscicauda Malloch, 1918: 269 - Washington,Oregon References: Malloch (1923)
= Phaonia fuscinervis Stein, 1920: 7
Phaonia harti Malloch, 1923: 266 - Alberta,Idaho, Arizona, Kans.to Michigan and Massachusetts, s. toVa.
Phaonia houghii (Stein, 1898: 177) - British Columbia toQue., s. to California,Oklahoma, andN.J. References: Malloch (1923)
= Aricia houghii Stein, 1898: 177
= Phaonia inculta Stein, 1920: 8
Phaonia hybrida (Schnabl, 1888: 396) - Alaska, Yukon Territory,Nunavut,Palaearctic References: Huckett (1965)
= Aricia hybrida Schnabl, 1888: 396
Phaonia imitatrix Malloch, 1919: 61c - USA (Alaska); Canada(Yukon) References: Huckett (1965)
Phaonia inenarrabilis Huckett, 1965: 314 - Nunavut,Alberta
Phaonia laticornis Malloch, 1923: 279 - Wisconsin, Michigan, Illinois, New Hampshire toN.C.
Phaonia lugubris (Meigen, 1826: 87) - Alaska, Yukon Territory, Nunavut, Greenland, cent. and n. Europe, e.Asia References: Huckett (1965) (as morio)
= Anthomyia lugubris Meigen, 1826: 87
= Aricia morio Boheman, 1844: 103 (nomen nudum)
= Aricia morio Zetterstedt, 1845: 1399
= Aricia plumbea Zetterstedt of Lundbeck, 1898: 308
Phaonia magnicornis (Zetterstedt, 1845: 1666) - Alaska, Yukon Territory,Palaearctic References: Huckett (1965)
= Anthomyza magnicornis Zetterstedt, 1845: 1666
= Aricia medisecta Pandellé, 1899: 102
Phaonia marylandica Malloch, 1923: 265 - Maine
Phaonia monticola Malloch, 1918: 266 - Alaska to Newfoundland, s. to Arizona andColo. References: Malloch (1923), Huckett (1965)
= Aricia morio Zetterstedt of Stein, 1920: 17
Phaonia neglecta Huckett, 1966: 299 - Alberta,Washington, Oregon,Colorado
Phaonia nigricans Johannsen, 1916: 395 - Alberta to South Dakota,Utah, Wisconsin to Quebec and Newfoundland, s. toD.C. References: Malloch (1923)
= Phaonia nigricans Johannsen, 1916: 395
= Spilogaster cayugae Johannsen, 1917: 327
= Phaonia nervosa Stein, 1920: 12
Phaonia nigricauda Malloch, 1918: 268 - California References: Malloch (1923)
Phaonia pallidisquama (Zetterstedt, 1849: 3288) - Nunavut (Baffin I.), Greenland, Quebec, n.w.Europe References: Huckett (1965)
= Aricia pallidisquama Zetterstedt, 1849: 3288
= Aricia anthracina Zetterstedt, 1860: 6227
Phaonia pallidosa Huckett, 1965: 907 - British Columbia and Idaho toCalif.
= Dialyta pallida Stein, 1920: 22
Phaonia pallidula Coquillett, 1902: 122 - Tex.to s. Ga.toMass References: Malloch (1923)
= Phaonia dulcis Stein, 1920: 5
Phaonia parviceps Malloch, 1918: 267 - Oregon References: Malloch (1923)
= Phaonia caesia Stein, 1920: 4
Phaonia peregrinans Huckett, 1965: 314 - Alaska
Phaonia perfida Stein, 1920: 13 - California,Utah References: Malloch (1923)
Phaonia picealis Huckett, 1965: 315 - Alaska
Phaonia pratensis (Robineau-Desvoidy, 1830: 485) - Michigan, Ontario, New Hampshire, Ohio,Europe References: Collin (1951), Pont (1984)
= Musca laeta (Fallén, 1823: 56) (tentative placement)
= Anthomyia trigonalis Meigen, 1826: 127
= Mydaea nigripes Robineau-Desvoidy, 1830: 481
= Euphemia pratensis Robineau-Desvoidy, 1830: 485
= Euphemia tibialis Robineau-Desvoidy, 1830: 486
= Anthomyia betuleti Bouché, 1834: 78
= Aricia brevivillosa Macquart, 1835: 292
= Aricia nigripes Schnabl, 1888: 454
= Aricia maculipennis Strøm, 1896: 238
= Phaonia laetabilis Collin, 1951: 2
Phaonia prisca Stein, 1920: 14 - New York References: Malloch (1923)
Phaonia protuberans Malloch, 1923: 247 - Washington to Colorado,Alberta to Labrador, s.toN.Y. References: Huckett (1965)
Phaonia proxima (Wulp, 1869: 85) - Alaska, Yukon Territory,Washington,Michigan References: Huckett (1965)
= Aricia proxima Wulp, 1869: 85
Phaonia pudoa Hall, 1937: 215 - Washington,Montana
Phaonia pulvillata (Stein, 1904: 422) - New York to Georgia References: Malloch (1923)
= Aricia pulvillataStein, 1904: 422
Phaonia quieta Stein, 1920: 14 - Oregon,California,Utah,Michigan References: Malloch (1923)
Phaonia reclusa Huckett, 1966: 300 - California
Phaonia reflecta Huckett, 1966: 301 - California
Phaonia reversa Huckett, 1966: 303 - California
Phaonia rufibasis Malloch, 1919: 207 - Alberta, Utah, Que.toMd References: Malloch (1923)
Phaonia rugia (Walker, 1849: 923) - Alaska to Labr., s. to Utah, Colo.andN.H. References: Malloch (1923) (as incerta), Huckett (1965)
= Anthomyia rugia Walker, 1849: 923
= Aricia brunneinervis Stein, 1898: 183
= Phaonia incerta Malloch, 1923: 250
Phaonia savonoskii Malloch, 1923: 248 - Nunavut,Washington,Alberta,Manitoba References: Huckett (1965)
Phaonia serva (Meigen, 1826: 86) - N.W.T.to Newfoundland, s. to Wyo. and Georgia,Palaearctic References: Malloch (1923), Huckett (1965)
= Anthomyia serva Meigen, 1826: 86
= Fellaea agilis Robineau-Desvoidy, 1830: 478
= Fellaea erratica Robineau-Desvoidy, 1830: 478
= Anthomyia interlatens Walker, 1853: 120
Phaonia sobriana Huckett, 1966: 304 - California
Phaonia soccata (Walker, 1849: 941) - Alberta, Ontario to MainetoN.Y. References: Malloch (1923), Huckett (1965)
= Anthomyia soccata Walker, 1849: 941
Phaonia striata (Stein, 1898: 179) - Washington,California References: Malloch (1923)
= Aricia striata Stein, 1898: 179
Phaonia subfusca Malloch, 1923: 273 - Arkansas, Ohio,New York
Phaonia subfuscinervis (Zetterstedt, 1838: 673) - Alaska to Greenland, s. to Manitoba and Quebec, New Hampshire,n.w.Europe References: Huckett (1965)
= Anthomyza inconspicua Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza subfuscinervis Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza turpis Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza vicina Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza inconspicua Zetterstedt, 1838: 673
= Anthomyza subfuscinervis Zetterstedt, 1838: 673
= Anthomyza turpis Zetterstedt, 1838: 673
Phaonia tenebriona Huckett, 1965: 315 - Yukon
Phaonia tipulivora Malloch, 1923: 252 - New York,New Hampshire, Maine,Massachusetts,Connecticut
Phaonia trivialis Malloch, 1923: 278 - Alberta
Phaonia uniseriata Malloch, 1923: 268 - Washington
Phaonia versicolor Stein, 1920: 16 - Colorado References: Malloch (1923)
Phaonia winnemanae Malloch, 1919: 3 - Wisconsin,Michigan, New York,Massachusetts, Maine,New Hampshire References: Malloch (1923)
= Phaonia apta Stein, 1920: 4

Genus Dolichophaonia Carvalho, 1993: 19 (2 species)
Dolichophaonia texensis (Malloch, 1923) - California, Arizona,Florida,Mexico
+ssp. flavofemorata Malloch - Texas, Florida
Dolichophaonia limbinervis (Stein, 1918: 208) - Mexico; North America References: Malloch (1923)

References:
de Carvalho, C. J. B. DOLICHOPHAONIA, GEN. N.(DIPTERA, MUSCIDAE, PHAONIINAE) DESCRIÇOES, NOVAS COMBINAÇOES, SINONIMIAS. Revista Brasiliera de Entomologia 37(1): 19-34.
de Carvalho, C.J.B. & Couri, M.S. 2002. Part I. Basal groups, pp. 17-132. In: Carvalho, C.J.B. de (ed.), Muscidae (Diptera) of the Neotropical Region: taxonomy. Iniversidade Federal do Rio de Janeiro, Curitiba. 293 pp.
de Carvalho, C.J.B., Couri, M.S., Pont, A.C., Pamplona, D. & Lopes, S.M. 2005. A catalogue of the Muscidae (Diptera) of the Neotropical Region. Zootaxa 860, 282 pp.
Collin, J.E. 1951. Phaonia laetabilis sp.n. with notes on some other related Anthomyiidae (Diptera). Entomologist’s Record and Journal of Variation 63(2): 1-5.
Couri, M.S. & de Carvalho, C.J.B. 2002. Part II. Apical groups, pp. 133-260. In: Carvalho, C.J.B. de (ed.), Muscidae (Diptera) of the Neotropical Region: taxonomy. Iniversidade Federal do Rio de Janeiro, Curitiba. 293 pp.
Falk, S.J, and Pont, A.C. 2017. A Provisional Assessment of the Status of Calypterate flies in the UK. Natural England. Commissioned Reports, Number 234.
Fallen, C.F. 1820-1825. Monographia Muscidum Sveciae.
Hall, D.G. 1937. New muscoid flies (Diptera) in the United States National Museum. Proceedings of the United States National Museum 84[3011]: 201-216.
Huckett, H.C. 1965. The Muscidae of Northern Canada, Alaska and Greenland (Diptera). Mem. ent. Soc. Can. 42, 369 pp.
Huckett, H.C. 1966. New species of Anthomyiidae and Muscidae from California (Diptera).Proceedings of the California Academy of Sciences 34: 235-305.
Huckett, H.C. 1973. The Anthomyiidae and Muscidae of Mt. Katahdin, Maine (Diptera). J. N.Y. ent. Soc. 80[1972]: 216-233.
Malloch, J.R. 1918. Diptera from the southwestern United States. Paper IV. Anthomyiidae. Transactions of the American Entomological Society 44: 263-319, 1 pl.
Malloch, J.R. 1919. One new genus and two new species of Anthomyiidae from the vicinity of Washington, D. C. (Diptera). Proceedings of the Biological Society of Washington 32: 1-14.
Malloch, J.R. 1919. Some new eastern Anthomyiidae (Diptera). Proceedings of the Biological Society of Washington 32: 207-210.
Malloch, J.R. 1919. The Diptera collected by the Canadian Expedition, 1913-1918 (Excluding the Tipulidae and Culicidae). Pp. 34c-90c. In: Anderson, R. M. (ed.), Report of the Canadian Arctic Expedition 1913-1918. Vol. 3: Insecta. 90 pp., 10 pls.
Malloch, J.R. 1920. Descriptions of new North American Anthomyiidae (Diptera). Transactions of the American Entomological Society 46: 133-196, 3 pls.
Malloch, J.R. 1923. Flies of the anthomyiid genus Phaonia and related genera. Transactions of the American Entomological Society 48[1922]: 227-282.
Meigen, J.W. 1826. Systematische Beschreibung der bekannten europäische n zweiflugeligen Insekten. Funfter Theil. Schulz-Wundermann, Hamm. xii + 412 pp.
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Stein, P. 1914. Versuch, die Gattungen und Arten unserer Anthomyiden nur nach dem weiblichen Geschlecht zu bestimmen, nebst Beschreibung einiger neuen Arten. Archiv für Naturgeschichte (Abt. A) 79 (8)[1913]: 4-55.
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Posted on 20 de junho de 2022, 07:25 AM by aispinsects aispinsects | 0 comentários | Deixar um comentário

22 de junho de 2022

Systema Dipterorum's Checklist of Helina (Diptera: Muscidae) north of Mexico and potentially

The genus Helina has been called the second largest genus in Muscidae., with well over 400 species in the world (Wang et al., 2004), It is also one of the difficult to recognize genera of muscids along with Phaonia and Mydaea. The biology of Helina species is poorly studied. Many revisionary works lack any mention at all on biology, but most species are probably detritivores as larvae, living under bark, in rotting wood, in tree holes, in vertebrate nests, and more.
No catalogue for Muscidae exists for the Nearctic region after Huckett in Stone et al. (1965). Since then, several changes could have occured according to Systema Dipterorum. A catalogue to Neotropical Muscidae was created by de Carvalho et al. (2005). Quadrularia is now a part of Helina, transferring annosa, "laetifica" (=evecta), "laetifica var. nivalis" (=flavisquama), and "punctata " Stein (=steini) to Helina. Helina laetifica is considered to be a synonym of H. evecta, but its variety known as nivalis is considered one of H. flavisquama by Systema Dipterorum, which conflicts with BugGuide. Helina steini is proposed by Pont (2013) for Helina puctata Stein. Helina flavisquama is also now considered the new name for Helina basalis Zetterstedt. Helina duplicata is now a synonym of the Holarctic species Helina reversio, the latter thus being an introduced species not recognized until recently. Helina troene var. fulviventris is treated as a full species distinct from troene. Eupogonomyia neoborealis is transferred to Helina. Helina platykarenos and polychaeta are described from California by Huckett (196). Helina punctata Robineau-Desvoidy is a synonym of Helina sexmaculata. Helina rothi became a synonym of H. subvittata. Ariciella flavicornis is transferred to Helina and its new name Helina rubripalpis is delivered by Pont (1972). Helina setifer is described by Huckett (1965). Helina snyderi is described from Wyoming by Steystkal (1966). Once again, these are all from the perspective of Systema Dipterorum and often these changes are not accepted among workers in reality
Additional species were included in this checklist that are not known north of Mexico, but are implied in some way by Systema Dipterorum that they could reach into north. The species are included if the range notes detail "NE: Mexico" (not NT: Mexico) with NE meaning Nearctic and NT meaning Neotropical or something more explicit. These species are Helina caneo (range notes: "(NE: NE) Mexico"), H. meraca (range notes: "(NT: NT NE) Mexico"), mulcata (range notes: "(NT: NT NE) Mexico; ?North America"), parvula (range notes: "(NE: NE PA NT AF) Mexico"), sera (its synonym, H. arroya has range notes: "NE"), and H. tarsalis (range notes: "(NE: NE) Mexico"). Helina depuncta and H. refusa were listed in Stone et al. (1965) with strong doubts as to their Nearctic occurences. There remains no reason to believe they occur in the Nearctic.
In order to key or identify these specimens, several keys are needed at hand: Wulp (1896), Malloch (1921), Snyder (1949b), Huckett (1965), and more. Original descriptions may also need to be consulted, especially for specimens from subtropical areas. Snyder (1949) is the major revision to use. In that paper, H. rothi is H. subvittata, H. anceps is nigribasis, and H. obscuripes is cothurnata, according to Systema Dipterorum.

Checklist (67 species):
Genus Helina Robineau-Desvoidy, 1830: 493
Helina abiens (Stein, 1898: 193) - Arizona, North Dakota to s. Quebec, s. toN.C. References: Snyder (1949b)
= Spilogaster abiens Stein, 1898: 193
Helina algonquina Malloch, 1922: 96 - North Carolina,Florida References: Snyder (1949b)
Helina annosa (Zetterstedt, 1838: 663) - Alaska to Labr.,s. to California and New York,Palaearctic References: Malloch (1921), Snyder (1949a)
= Anthomyza annosa Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza annosa Zetterstedt, 1838: 663
= Aricia bicolor Pokorny, 1889: 549
= Aricia multisetosa Strobl, 1898: 238
Helina barpana (Walker, 1849: 933) - Nova Scotia
= Anthomyia barpana Walker, 1849: 933
Helina bicolorata (Malloch in Cole and Lovett, 1919: 253) - British Columbia, Washington,California References: Snyder (1949b)
= Aricia bicolorata Malloch in Cole and Lovett, 1919: 253
= Mydaea aperta Stein, 1920: 28
Helina bispinosa Malloch, 1920: 142 - British Columbia toMan., s. toCalif. and New Mexico References: Snyder (1949b)
Helina bohemani (Ringdahl, 1916: 235) - British Columbia to Manitoba, s.to California and New Mexico, also Illinois, n. Europe,Russia References: Snyder (1949b)
= Helina subeiensis Ma & Wang, 1992: 407
= Mydaea bohemani Ringdahl, 1916: 235
Helina canadensis Snyder, 1949b: 136 - Quebec,New York
Helina caneo Snyder, 1941: 10 - Mexico
Helina cinerella (Wulp, 1867: 150) - Alaska to Labr., s. to Calif.and Michigan,Palaearctic References: Snyder (1949b) (as vanderwulpi)
= Aricia cinerella Wulp, 1867: 150
= Aricia vanderwulpi Schnabl, 1888: 387
= Aricia propinqua Strøm, 1896: 239
= Aricia brevis Stein, 1898: 180
= Aricia menechma Pandellé, 1898: 56
= Helina aldrichi Snyder, 1949: 122
= Helina tuleskovi Lavciev, 1968: 63
Helina consimilata Malloch, 1920: 144 - Washington, Colorado,Wisconsin to New Hampshire andMass. References: Snyder (1949b)
Helina cothurnata (Rondani, 1866: 116) - Alaska to N.W.T., s. to California and New Hampshire, Palaearctic References: Snyder (1949b) (as obscuripes)
= Spilogaster cothurnata Rondani, 1866: 116
= Anthomyza obscuripes Zetterstedt of Stein, 1916: 65
Helina cruciata Snyder, 1941: 9 - Arizona References: Snyder (1949b)
Helina evecta (Harris, 1780: 125) - Alaska to Nfld., s. to California and Tennessee, cent. Amer.,Palaearctic References: Malloch (1921), Snyder (1949a)
= Musca evectus Harris, 1780: 125
= Musca lucorum Fallén, 1823: 55
= Mydina laetifica Robineau-Desvoidy, 1830: 500
= Mydina soror Robineau-Desvoidy, 1830: 501
= Anthomyia pylone Walker, 1849: 928
=Anthomyia incerta Walker, 1853: 354
= Anthomyia solita Walker, 1853: 354
= Spilogaster venosa Rondani, 1877: 98
= Aricia graefenbergiana Schnabl, 1888: 447
= Aricia mohyleviensis Schnabl, 1888: 449
= Aricia obscurataeformis Schnabl, 1888: 383
= Hyetodesia abacta Giglio-Tos, 1893: 8
= Hyetodesia abdicta Giglio-Tos, 1893: 8
= Mydaea vulnifera Villeneuve, 1927: 266
= Spilogaster limbovenosa Hennig, 1961: 228
Helina exilis (Stein, 1920: 30) - Wisconsin, Michigan,New York,Quebec,Connecticut References: Snyder (1949b)
= Mydaea exilis Stein, 1920: 30
Helina flavisquama (Zetterstedt, 1849: 3287) - Alaska to Nunavut,s. to California and Labrador,Newfoundland,Palaearctic. References: Snyder (1949b)
= Anthomyza basalis Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza basalis Zetterstedt, 1838: 663
= Anthomyza nivalis Zetterstedt, 1838: 663
= Aricia flavisquama Zetterstedt, 1849: 3287
= Aricia setigera Stein, 1900: 305
= Mydaea setitibia Stein, 1914: 19
= Mydaea flavocalyptrata Stein, 1920: 31
= Helina hylemyioides Malloch, 1920: 137
Helina floridensis Snyder, 1949: 126 - Florida References: Snyder (1949b)
Helina fulvisquama (Zetterstedt, 1845: 1491) - Alaska to Labrador, s. to California and New Hampshire,Lapland, n.Europe,Russia References: Snyder (1949b)
= Aricia fulvisquama Zetterstedt, 1845: 1491
= Phaonia arctica Schnabl & Becker, 1915: 50
= Helina tuberculata Malloch, 1919: 277
Helina fulviventris (Bigot, 1885: 291) - B.C.s. to Calif. References: Snyder (1949b) (as synonym of troene)
= Spilogaster fulviventris Bigot, 1885: 291
= Mydaea varia Stein, 1920: 36
= Helina troene var. fulviventris Bigot of Stone et al., 1965
Helina garretti Snyder, 1949: 140 - British Columbia
Helina griseogaster Snyder, 1949: 138 = Colorado
Helina humilis (Stein, 1920: 32) - British Columbia to California, Arizona, Wyo.,andColo. References: Snyder (1949b)
= Mydaea humilis Stein, 1920: 32
Helina johnsoni Malloch, 1920: 141 - Wisconsin,Quebec and N.S., s.toGa. References: Snyder (1949b)
Helina keremeosa Snyder, 1949: 156 - California References: Snyder (1949b)
Helina lasiosterna Snyder, 1941: 9 - Mexico; North America References: Snyder (1949b)
Helina laxifrons (Zetterstedt, 1860: 6200) - Alaska to Que., s. to Utah and Georgia,Palaearctic References: Snyder (1949b)
= Aricia laxifrons Zetterstedt, 1860: 6200
= Aricia curvipes Schnabl, 1888: 383
= Aricia nigripennis Schnabl, 1888: 378
= Spilogaster nigricans Stein, 1898: 198
= Mydaea tinctipennis Stein, 1916: 69
Helina linearis Malloch, 1920: 139 - Washington, Alberta, Montana,Minnesota to Quebec, s. toIll. References: Snyder (1949b)
Helina longicornis (Zetterstedt, 1838: 678) - Yukon Territory, Nunavut,Manitoba,Palaearctic References: Snyder (1949b) (notes), Huckett (1965)
= Anthomyza longicornis Zetterstedt, 1837: 44 (nomen nudum)
= Anthomyza longicornis Zetterstedt, 1838: 678
= Spilogaster angulicornis Pokorny, 1889: 555
Helina luteisquama (Zetterstedt, 1845: 1492) - Alaskato Nunavut, s. to Manitoba and Labrador,Palaearctic References: Snyder (1949b)
= Aricia luteisquama Zetterstedt, 1845: 1492
Helina maculipennis (Zetterstedt, 1845: 1475) - Alaska, British Columbia and Alberta, s. to California and New Mexico, alsoMan.,Labrador, Brazil, Europe, Asia References: Snyder (1949b) (as neopoeciloptera)
= Helina neopoeciloptera Malloch, 1920: 139
= Helina poeciloptera Malloch, 1918: 271
Helina marguerita Snyder, 1949: 137 - Yukon Territory, Nunavut, California,Quebec, Ontario,Labrador References: Snyder (1949b)
Helina meraca (Wulp, 1896: 325) - Mexico References: Wulp (1896) (as Limnophora), Pont (1972) (not easily accessible), de Carvalho et al. (2005)
= Limnophora meraca Wulp, 1896: 325
= Helina meraca>/i> Wulp of Pont, 1972 (new combination)
Helina mulcata
(Giglio-Tos, 1893: 7) - Mexico; ?North America References: Wulp (1896) (as Hyetodesia), Stein (1918) (notes) (as Mydaea), de Carvalho et al. (2005)
= Hyetodesia mulcata Giglio-Tos, 1893: 7
Helina multiseriata Malloch, 1922: 95 - British Columbia and Alberta,s. to California andColo. References: Snyder (1949b)
Helina neoborealis Snyder, 1949: 122 - Alaska, Nunavut (Victoria and Ellesmere Is.),California,Colorado References: Huckett (1965) (as Eupogonomyia)
= Eupogonomyia neoborealis Snyder of Stone et al., 1965
Helina nigribasis Malloch, 1920: 143 - Alaska to Arizona, Wisconsin & Illinois to Maryland and Labrador, s. to Florida. References: Snyder (1949b)
= Anthomyza anceps Zetterstedt of Stein, 1904: 449
Helina nigripennis (Walker, 1849: 929) - Alaska to California and Colorado, also Manitoba to Newfoundland, s.toMich. References: Snyder (1949b)
= Anthomyia nigripennis Walker, 1849: 929
= Aricia nitida Stein, 1898: 185
= Spilogaster crepuscularis Stein, 1898: 201
Helina nigrita Malloch, 1920: 139 - Alberta toMan., s. toCalif. and New Mexico References: Snyder (1949b)
Helina nudibasis Snyder, 1949: 147 - New Mexico
Helina obscurata (Meigen, 1826: 89) - Alaska to Labrador,s. to California, Illinois, and New York,Palaearctic References: Snyder (1949b)
= Anthomyia obscurata Meigen, 1826: 89
= Mydina fuliginosa Robineau-Desvoidy, 1830: 498
= Mydina vernalis Robineau-Desvoidy, 1830: 498
= Anthomyza sahlbergi Zetterstedt, 1837: 43 (nomen nudum)
= Anthomyza sahlbergi Zetterstedt, 1838: 664
= Anthomyza veterana Zetterstedt, 1838: 662
= Aricia duplaris Zetterstedt, 1845: 1411
= Aricia sordidiventris Zetterstedt, 1845: 1416
= Anthomyia detracta Walker, 1853: 122
= Helina nasoni Malloch, 1920: 138
Helina obscuratoides (Schnabl, 1887: 347) - Alberta, Manitoba,South Dakota,Palaearctic References: Snyder (1949b)
= Aricia obscuratidea Pandellé, 1898: 59
= Aricia obscuratoides Schnabl, 1887: 347
Helina obscurinervis (Stein, 1898: 199) - Oregon, California, Kansas to Quebec, s. to Texas andFla. References: Snyder (1949b)
= Spilogaster obscurinervis Stein, 1898: 199
Helina orbitaseta (Stein, 1898: 186) - Wash.to California andAriz., alsoMo. References: Snyder (1949b)
= Aricia orbitaseta/i> Stein, 1898: 186
Helina oregonensis
(Malloch in Cole and Lovett, 1919: 254) - British Columbia,Idaho,Utah References: Snyder (1949b) (notes)
= Aricia oregonensis Malloch in Cole and Lovett, 1919: 254
Helina parvula (Wulp, 1896: 321) - Mexico References: Snyder (1941), de Carvalho et al. (2005)
= Spilogaster parvula Wulp, 1896: 321
Helina pectinata (Johannsen, 1916: 392) - British Columbia to Manitoba, s. toCalif. and Arkansas, also Que.to N.S. toN.Y. References: Snyder (1949b)
= Mydaea pectinata Johannsen, 1916: 392
= Mydaea biseriata Stein, 1920: 29
= Helina mimetica Malloch, 1920: 142
Helina platykarenos Huckett, 1966: 286 - California
Helina polychaeta Huckett, 1966: 288 - California
Helina procedens (Walker, 1861: 315) - Mexico; North America References: Snyder (1949b)
= Aricia procedens Walker, 1861: 315
= Spilogaster uniseta Stein, 1898: 192
Helina reversio (Harris, 1780: 146) - Alaska to Nfld., s. to California and New York,Palaearctic References: Malloch (1921) (as duplicata), Huckett (1965) (as duplicata)
= Musca reversio Harris, 1780: 146
= Anthomyia compuncta Wiedemann, 1817: 80
= Anthomyia duplicata Meigen, 1826: 92
= Helina tibialis Robineau-Desvoidy, 1830: 494
= Mydina claripennis Robineau-Desvoidy, 1830: 497
= Mydina communis Robineau-Desvoidy, 1830: 497
= Mydina limpidipennis Robineau-Desvoidy, 1830: 498
= Mydina nigricans Robineau-Desvoidy, 1830: 497
= Mydina nigripes Robineau-Desvoidy, 1830: 498
= Anthomyia quadripunctata Brullé, 1833: 315
= Spilogaster caesia Macquart, 1835: 296
= Spilogaster quadrivittata Macquart, 1844: 163
= Anthomyia decedens Walker, 1853: 121
= Anthomyia fixa Walker, 1853: 121
= Anthomyia infixa Walker, 1853: 132
= Anthomyza vilis Zetterstedt, 1845: 1669
= Anthomyza flavogrisea Zetterstedt, 1860: 6275
= Aricia prospinosa Pandellé, 1898: 56
= Mydaea multimaculata Schnabl, 1911: 95
= Mydaea tagojana Santos Abreu, 1976: 115
Helina rubripalpis (Wulp, 1896: 320) - Mexico, Costa Rica; Texas References: Wulp (1896) (as Spilogaster), Stein (1920) (as Mydaea), Pont (1972), de Carvalho et al. (2005)
= Spilogaster rubripalpis Wulp, 1896: 320
= Ariciella flavicornis Malloch, 1918: 66
= Helina flavicornis Malloch, 1918: 66
Helina rufitibia (Stein, 1898: 181) - Alberta to Quebec, s. to California andGa. References: Snyder (1949b)
= Aricia rufitibia Stein, 1898: 181
= Hebecnema rufitibia Stein, 1908: 95
= Helina rufitibialis Hennig, 1957: 152
= Hebecnema luteoligaster Santos Abreu, 1976: 75
= Hebecnema nitidiuscula Santos Abreu, 1976: 76
= Hebecnema thoracica Santos Abreu, 1976: 76
Helina sera (Giglio-Tos, 1893: 9) - Mexico References: Snyder (1949c), de Carvalho et al. (2005)
= Spilogaster sera Giglio-Tos, 1893: 9
= Helina arroya Snyder, 1949: 6
Helina setifer Huckett, 1965: 256 - Yukon Territory
Helina sexmaculata (Preyssler, 1791: 88) - South Dakota and Minnesota, s. to Colorado and Missouri, Massachusetts to Georgia,Palaearctic, New Zealand References: Snyder (1949b) (as uliginosa)
= Musca deduco Harris, 1780: 125
= Musca sexmaculata Preyssler, 1791: 88
= Musca uliginosa Fallén, 1825: 81
= Rohrella punctata Robineau-Desvoidy, 1830: 492
= Anthomyza flavicoxa Zetterstedt, 1860: 6277
Helina spinicosta (Zetterstedt, 1845: 1641) - Alaska,Nunavut, n. Europe, n.USAS.R. References: Snyder (1949b) (notes), Huckett (1965)
= Anthomyza spinicosta Zetterstedt, 1845: 1641
= Anthomyza congenulata Zetterstedt, 1860: 6266
= Ecliponeura spinicosta Emden, 1941: 257
= Helina spinicostata Pont, 1980: 736
Helina spinilamellata Malloch, 1920: 140 - Montana References: Snyder (1949b)
Helina spinosa (Walker, 1849: 926) - Alaska to Labrador, s. toCalif. and New Mexico References: Snyder (1949b)
= Anthomyia spinosa Walker, 1849: 926
= Aricia latifrontata Malloch, 1918: 270
Helina spuria Malloch, 1920: 144 - Oregon, Colorado References: Snyder (1949b)
Helina squalens (Zetterstedt, 1838: 669) - Alaska to Labr., s. to Manitoba, n.w.Europe References: Snyder (1949b)
= Anthomyza borealis Zetterstedt, 1837: 44
= Anthomyza squalens Zetterstedt, 1837: 43
= Anthomyza borealis Zetterstedt, 1838: 679
= Anthomyza squalens Zetterstedt, 1838: 669
Helina snyderi Steyskal, 1966: 177 - Wyoming
Helina steini Pont, 2013
= Aricia punctata Stein, 1898: 182
= Quadrularia punctata Stein of Stone et al., 1965
Helina subvittata (Séguy, 1923: 336) - Europe: (Andorra), Russia: Asia: Japan; Alaska to Labrador, s. to California and New York,Palaearctic References: Snyder (1949b) (as rothi)
= Helina rothi Ringdahl, 1939: 150
= Phaonia subvittata Séguy, 1923: 336
Helina tarsalis (Stein, 1918: 220) - Mexico References: Stein (1920) (as Mydaea), Pont (1972) (not easily accessible), de Carvalho et al. (2005)
= Mydaea tarsalis Stein, 1918: 220
= Helina tarsalis Stein of Pont, 1972 (new combination)
Helina toga Snyder, 1949: 158 - Michigan
Helina troene (Walker, 1849: 936) - British Columbia to Newfoundland, s. to Calif.andGa. References: Snyder (1949b)
= Anthomyia lysinoe Walker, 1849: 938
= Anthomyia troene Walker, 1849: 936
= Spilogaster amoeba Stein, 1898: 190
= Spilogaster pubiceps Stein, 1898: 194
Helina ute Snyder, 1949: 155 - California
Helina villihumilis Snyder, 1949: 156 - British Columbia

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